Effects of ipsilateral and contralateral precursors on overshoot.

Overshoot is defined as the decrease in threshold as a brief signal is moved from the beginning to near the temporal center of a longer duration, broadband noise masker. Overshoot can be reduced when another noise (a precursor) is presented just prior to the masker. The purpose of the present investigation was to follow up on a recent psychophysical study which showed that overshoot could be reduced by a precursor presented to the ear contralateral to that receiving the masker and signal. The signal was a 20-ms, 4000-Hz tone that was presented at the beginning or in the temporal center of a 400-ms broadband noise masker. In the first experiment, a 200-ms broadband precursor was presented either to the ipsilateral or to the contralateral ear. The ipsilateral precursor reduced overshoot for all ten subjects, but the contralateral precursor reduced overshoot for only four of the ten subjects. In a supplementary experiment, the contralateral precursor failed to reduce overshoot in a new group of five subjects, both when tested with supra-aural headphones and with insert earphones. In the second experiment, the four subjects who showed an effect of the contralateral precursor in experiment 1 were tested under conditions where the bandwidth of the precursor was manipulated, resulting in either a narrow-band precursor centered at 4000 Hz, a low-band precursor with energy primarily below 4000 Hz, or a high-band precursor with energy primarily above 4000 Hz. There was a tendency for the effectiveness of the ipsilateral and contralateral precursors to be affected similarly (though to different degrees) by changes in the spectral content of the precursor. These results suggest that the effect of the contralateral precursor is not due to a timing cue, and that the processing underlying the effectiveness of ipsilateral and contralateral precursors may be largely the same.

Contributions of suppression and excitation to simultaneous masking: effects of signal frequency and masker-signal frequency relation.

This study investigated the contributions of suppression and excitation to simultaneous masking for a range of masker frequencies both below and above three different signal frequencies (750, 2000, and 4850 Hz). A two-stage experiment was employed. In stage I, the level of each off-frequency simultaneous masker necessary to mask a signal at 10 or 30 dB sensation level was determined. In stage II, three different forward-masking conditions were tested: (1) an on-frequency condition, in which the signals in stage I were used to mask probes of the same frequency; (2) an off-frequency condition, in which the off-frequency maskers (at the levels determined in stage I) were used to mask the probes; and (3) a combined condition, in which the on- and off-frequency maskers were combined to mask the probes. If the off-frequency maskers simultaneously masked the signal via spread of excitation in stage I, then the off-frequency and combined maskers should produce considerable forward masking in stage II. If, on the other hand, they masked via suppression, they should produce little or no forward masking. The contribution of suppression was found to increase with increasing signal frequency; it was absent at 750 Hz, but dominant at 4850 Hz. These results have implications for excitation pattern analyses and are consistent with stronger nonlinear processing at high rather than at low frequencies.

Effects of ipsilateral and contralateral precursors on the temporal effect in simultaneous masking with pure tones.

In tone-on-tone masking, thresholds often decrease as the onset of the signal is delayed relative to the onset of the masker, especially when the frequency of the masker is higher than the frequency of the signal. This temporal effect was studied here by using a tonal "precursor," whose offset preceded the onset of the tonal masker (and signal). Under the right conditions, the precursor can reduce or eliminate the temporal effect by decreasing the threshold for a signal at masker onset, presumably for the same reason that the threshold decreases as a signal is delayed relative to the onset of a masker. In the present study, the frequency of the signal was 4000 Hz, and the frequency of the masker and precursor was typically 5000 Hz. In experiment 1, the precursor was presented to the ear receiving the masker and signal (ipsilateral precursor); in experiment 2, it was presented to the opposite ear (contralateral precursor). The results from experiment 1 can be summarized as follows: the ipsilateral precursor (a) reaches its maximum effectiveness (in reducing the temporal effect) for precursor durations of 200-400 ms; (b) is ineffective once the delay between its offset and the onset of the masker reaches about 50-100 ms; (c) is generally ineffective when its level is 10 or more dB lower than the level of the masker, but is effective when its level is equal to or greater than the level of the masker; and (d) becomes progressively less effective as its frequency is either increased or decreased relative to the frequency of the masker. The results from experiment 2 can be summarized simply by stating that the contralateral precursor is ineffective in reducing the temporal effect. These results suggest that the effect of the precursor may be mediated peripherally.

Temporal integration in the presence of off-frequency maskers.

Temporal integration was measured at a relatively low and a relatively high signal frequency under conditions of off-frequency masking. The masker was typically gated for 300 ms, and the signal was presented 70 ms after masker onset. In experiment 1, the signal frequency was 500 or 2000 Hz. Temporal integration was measured in quiet and in the presence of a masker whose frequency was lower or higher than the signal frequency. In all listening situations, there was less integration at 2000 Hz than at 500 Hz. This effect of frequency was particularly dramatic in the presence of a lower frequency masker, where there was almost no integration at 2000 Hz. Experiment 2 showed that this dramatic effect of frequency cannot be understood in terms of the underlying psychometric functions. Experiment 3 measured temporal integration at 750 and 2000 Hz for a large number of masker-signal frequency separations for both a tonal and a noise masker, and in conditions where the masker was gated or continuous. The results with the gated tonal masker largely confirmed the results of experiment 1. The results with the continuous tonal masker and the gated or continuous noise masker, however, were quite different. In those cases, the amount of temporal integration at both signal frequencies was more or less independent of the masker-signal separation; the masked temporal integration was nearly equal to the integration in quiet. Thus based on the conditions evaluated here, off-frequency masked temporal integration differs substantially from integration in quiet only for gated tonal maskers located considerably lower in frequency than the signal. It is unclear how to account for this finding, although it may be related to attentional factors.

Effects of aspirin on psychophysical measures of frequency selectivity, two-tone suppression, and growth of masking.

Three psychophysical measures of nonlinearity were evaluated before and during a course of aspirin ingestion to investigate the role of outer hair cells (OHCs) in these measures, as aspirin is thought to alter the functioning of OHCs. Six normal-hearing individuals received a moderate dose (3.9 g/day) of aspirin for four days, producing essentially flat, temporary hearing losses that ranged from 5-20 dB. The losses were about 2 dB greater for a 300-ms signal than for a 15-ms signal, indicating reduced temporal integration with aspirin. On the final three days of aspirin use, three experiments were completed; each was designed to measure one aspect of nonlinear behavior: (1) the effects of level on frequency selectivity in simultaneous masking using notched-noise maskers, (2) two-tone suppression using forward maskers at the signal frequency (fs) and suppressor tones above fs, and (3) growth-of-masking functions in forward masking using a masking tone below fs. Signal frequencies of 750 and 3000 Hz were used to evaluate the effects of aspirin at relatively low- and high-frequency regions of the cochlea. In experiment 1, aspirin broadened the auditory filters and reduced the effect of level on frequency selectivity. In experiment 2, aspirin reduced or eliminated two-tone suppression. And, in experiment 3, aspirin reduced the slopes of the growth-of-masking functions. Thus, the aspirin was effective in reducing nonlinearity in all three experiments, suggesting that these measures reflect the same (or a similar) active, nonlinear mechanism, namely the compressive nonlinearity provided by the OHCs. In all experiments, aspirin tended to have larger detrimental effects on the nonlinear measures at 3000 Hz than at 750 Hz, which can be explained in terms of greater involvement of nonlinear processing at higher frequencies. Finally, these effects of aspirin were found to be similar to those observed in preliminary measurements in two subjects with mild, permanent hearing loss.

Growth of simultaneous masking for fm < fs: effects of overall frequency and level.

Growth-of-masking (GOM) functions were obtained in three groups of normal-hearing subjects using a simultaneous-masking paradigm. In all cases, the signal frequency (fs) was higher than the masker frequency (fm), either by a certain ratio (1.44) or by a certain distance [3 equivalent rectangular bandwidths (ERBs)]. The purpose was to evaluate the effect of overall frequency on the slope of the steep portion of the GOM function, and to evaluate the change in slope that can occur at high levels. Signal frequency ranged from 400 to 5000 Hz, and masker level ranged from 40 to 95 dB SPL. On average, the slope of the steep portion of the GOM function was about 1.4 for signal frequencies from 400 to 750 Hz, and 2.0 for signal frequencies from 1944 to 5000 Hz. This is consistent with the possibility that the cochlea may behave more linearly at the apical (low-frequency) region than at the basal (high-frequency) region. In addition, for signal frequencies at and above 750 Hz, the slope of the masking function changed from a value much greater than 1.0 to a value of 1.0 at high levels. The change in slope was better correlated with signal sensation level (i.e., amount of masking) than with either signal or masker SPL: the lack of a change at the lower signal frequencies may reflect the smaller amounts of masking there. The change to a linear growth of masking may represent a change in the response to the signal from compressive to linear at high levels.

Some effects of background noise on modulation detection interference.

Modulation thresholds were obtained for a 2000-Hz signal carrier modulated at a rate of 10 Hz. Thresholds were obtained without a masker carrier and in the presence of a masker carrier that was either unmodulated or modulated at a rate of 10 Hz and a depth of 100% (m(m) = 1.0). Of primary interest was whether the amount of interference caused by the masker was influenced by the frequency proximity of the masker to the signal, and whether background noise had an influence on that proximity effect. In general, for masker carriers higher in frequency than the 2000-Hz signal carrier, there was a tendency for the interference to decline as the masker was moved farther away from the signal for masker carriers lower than 2000 Hz, there was little or no proximity effect. Broadband noise eliminated the proximity effect obtained with an unmodulated masker, but not that obtained with a modulated masker. Results with a narrowband noise suggest that the broadband noise has its effect by masking the high-frequency side of the signal's excitation pattern. These results, as well as the results of an excitation pattern analysis, suggest that the proximity effect with all unmodulated masker may be mediated via a peripheral, within-channel interaction, whereas that with a modulated masker may be mediated via a central, across-channel interaction.

Psychophysical measures of auditory nonlinearities as a function of frequency in individuals with normal hearing.

In order to gain a better understanding of how auditory nonlinear phenomena vary as a function of location along the cochlea, several psychophysical measures of nonlinearity were examined as a function of signal frequency. Six normal-hearing individuals completed three experiments, each designed to measure one aspect of nonlinear behavior: (1) the effects of level on frequency selectivity in simultaneous masking, measured using notched-noise maskers at spectrum levels of 30 and 50 dB, (2) two-tone suppression, measured using forward maskers at the signal frequency (fs) and suppressor tones above fs, and (3) growth of masking, measured using forward maskers below fs at a signal/masker frequency ratio of 1.44. Four signal frequencies (375, 750, 1500, and 3000 Hz) were tested to sample the nonlinear behavior at different locations along the basilar membrane, in order to test the hypothesis that the apical (low-frequency) region of the cochlea behaves more linearly than the basal (high-frequency) region. In general, all three measures revealed a progressive increase in nonlinear behavior as signal frequency increased, with little or no nonlinearity at the lowest frequency, consistent with the hypothesis.

Psychophysical suppression as a function of signal frequency: noise and tonal maskers.

Physiological studies have suggested that the basal region of the cochlea is more nonlinear than the apical region. To evaluate this possibility psychophysically, suppression was investigated across signal frequency (250, 500, 1000, 2000, and 4000 Hz) in a forward-masking paradigm using both noise and tonal maskers/suppressors. Masker duration was 200 ms, signal duration was 20 or 40 ms, and signal delay was 0 or 20 ms; the longer delay was necessary to eliminate potential confusion effects observed with the (narrow-band) noise masker. When using a noise masker (spectrum level of 40 dB), suppression was determined by comparing the threshold in the presence of a broadband masker with that in the presence of a critical band (ERB) masker. When using a tonal masker (masker level of 50 dB SPL, suppressor level of 70 dB SPL, with the suppressor frequency being 1.2 times the masker/signal frequency), suppression was determined by comparing the threshold in the presence of the masker plus suppressor with that in the presence of the masker alone. The magnitude of suppression was determined either by the measured change in signal threshold or by the inferred change in masker level. Regardless of how suppression was quantified, for both masker types, the amount of suppression increased as signal frequency increased up to about 1000 Hz, but then reached an asymptote or decreased somewhat as signal frequency increased to 4000 Hz. The magnitude of suppression was much larger with a noise masker than with a tonal masker, which could be a result of the different number of components in the masker which might serve as a suppressor.

The effects of hearing loss and noise masking on the masking release for speech in temporally complex backgrounds.

Speech recognition was measured in three groups of listeners: those with sensorineural hearing loss of (presumably) cochlear origin (HL), those with normal hearing (NH), and those with normal hearing who listened in the presence of a spectrally shaped noise that elevated their pure-tone thresholds to match those of individual listeners in the HL group (NM). Performance was measured in four backgrounds that differed only in their temporal envelope: steady-state (SS) speech-shaped noise, speech-shaped noise modulated by the envelope of multi-talker babble (MT), speech-shaped noise modulated by the envelope of single-talker speech (ST), and speech-shaped noise modulated by a 10-Hz square wave (SQ). Threshold signal-to-noise ratios (SNRs) were typically best in the ST and especially the SQ conditions, indicating a masking release in those modulated backgrounds. SNRs in the SS and MT conditions were essentially identical to one another. The masking release was largest in the listeners in the NH group, and it tended to decrease as hearing loss increased. In 5 of the 11 listeners in the HL group, the masking release was nearly identical to that obtained in the NM group matched to those listeners; in the other 6 listeners, the release was smaller than that in the NM group. The reduced masking release was simulated best in those HL listeners for whom the masking release was relatively large. These results suggest that reduced masking release for speech in listeners with sensorineural hearing loss can only sometimes be accounted for entirely by reduced audibility.

Masking by sinusoidally amplitude-modulated tonal maskers.

In experiment 1, masking patterns were obtained with a tonal masker that was sinusoidally amplitude modulated (SAM) at a rate of 8 Hz and a depth (m) of 1.0. The signal was centered at a masker peak or masker valley. Masker frequency (fm) was 750, 1350, or 2430 Hz, and signal frequency (fs) ranged from 0.8 to 1.62 fm. Thresholds were generally higher for a signal in a masker peak than in a masker valley. The magnitude of this peak-to-valley (PV) difference was governed by fs/fm, rather than by fs, and was largest for fs > fm. The PV differences were smallest at the lowest fm, at least when fs > fm. In experiment 2, growth-of-masking functions were measured (fm = 1350 Hz, fs = 1.44fm). The masker was modulated at a depth (m) of 1.0, 0.75, or 0.50. These thresholds were compared with those obtained with an unmodulated masker in forward or simultaneous masking. The comparisons suggest that thresholds for a signal at a peak of an 8-Hz SAM masker are due to simultaneous masking, while those in a valley are due primarily to forward masking when m = 1.0 or simultaneous masking when m = 0.75 or 0.50. For these masker depths, the PV difference first increased but then decreased as masker level increased from 60 to 90 dB SPL. This was a consequence of the slope of the masking function for peak placement changing from a value greater than 2.0 to a value of 1.0 at the highest signal levels (an effect that was also observed with the unmodulated simultaneous masker), a result that may be understood in terms of basilar membrane nonlinearity.

The modulated-unmodulated difference: effects of signal frequency and masker modulation depth.

The masked threshold for a signal is often times lower when the masker is modulated than when it is unmodulated. The difference in masked thresholds is referred to as the modulated-unmodulated difference, or MUD. The purpose of the present study was to follow up on the results of a previous study [Bacon et al., J. Acoust. Soc. Am. 101, 1600-1610 (1997)] which showed that the MUD is larger for high than for low signal frequencies, both when the masker is no wider than a critical band (and the processing is solely within channel) and when it is broadband (and the processing may be both within and across channel). The present results indicate that the effects of signal frequency primarily exist only when the modulated masker is modulated at a depth greater than about 0.75, and that at these large depths, thresholds in the presence of the modulated masker are governed largely by forward masking. By far, the effect of signal frequency is larger with the broadband masker than with the critical-band masker, suggesting that there may be an across-channel process whose contribution is greater at high than at low signal frequencies. It is argued here that this across-channel process may be related to psychophysical suppression.

Amplitude modulation depth discrimination of a sinusoidal carrier: effect of stimulus duration.

Discrimination of the change in depth of sinusoidal amplitude modulation (AM) was investigated as a function of stimulus duration. The carrier frequency was 4000 Hz, the standard modulation depth (m) was either 0.1, 0.18, or 0.3, and the modulation rate was either 10, 20, 40, or 80 Hz. For all standard depths and modulation rates, threshold (delta m) decreased by more than a factor o two as stimulus duration doubled from the shortest duration used up to a certain duration (critical duration), beyond which the threshold decreased only slightly or remained constant. The critical duration corresponded to about four cycles of modulation. Psychometric functions were measured for different stimulus durations to examine the extent to which a multiple-looks model could explain the present data. This model provided a reasonable prediction of the change in AM depth discrimination threshold as a function of stimulus duration.

Effect of low-frequency gain reduction on speech recognition and its relation to upward spread of masking.

Speech recognition was measured in listeners with normal hearing and in listeners with sensorineural hearing loss under conditions that simulated hearing aid processing in a low-pass and speech-shaped background noise. Differing amounts of low-frequency gain reduction were applied during a high-frequency monosyllable test and a sentence level test to simulate the frequency responses of some commercial hearing aids. The results showed an improvement in speech recognition with low-frequency gain reduction in the low-pass noise, but not in the speech-shaped background noise. Masking patterns also were obtained with the two background noises at 70 and 80 dB SPL to compare with the speech results. There was no correlation observed between the masking results and the improvement in speech recognition with low-frequency gain reduction.

Masking by modulated and unmodulated noise: effects of bandwidth, modulation rate, signal frequency, and masker level.

The threshold for a sinusoidal signal masked by a band of noise is often times lower when the masking noise is modulated than when it is unmodulated. The difference in masked thresholds is referred to as the modulated-unmodulated difference, or MUD. These present experiments examined the effects of masker bandwidth, masker rate, and masker level on the MUD at several different signal frequencies. The MUD generally increased with increasing masker bandwidth; for masker bandwidths wider than a critical band (or an equivalent rectangular bandwidth-ERB), the results may be influenced by across-channel processes underlying comodulation masking release. The MUD for an ERB masker (MUDERB) was influenced less by masker rate than was the MUD for a broadband (BB) masker (MUDBB). The MUDERB and especially the MUDBB increased significantly with increasing masker level when the modulated masker was modulated at a depth (m) of 1.0, but not when it was modulated at a depth of 0.75. These results have significant implications for extending the MUD paradigm to hearing-impaired subjects. Finally, the MUDERB and the MUDBB increased with increasing signal frequency. This effect for the ERB masker is largely (if not completely) due to the wider absolute bandwidths at higher frequencies. The effect with the BB masker may be influenced by differences in the magnitude of suppression across frequency.

The effect of level and relative frequency region on the recovery of overshoot.

Overshoot--in particular, threshold for a signal near masker onset--can be reduced by presenting a stimulus (precursor) just prior to masker onset. The recovery of overshoot can be examined by varying the delay between the offset of the precursor and the onset of the masker, where "recovery" denotes an increase in the threshold for a signal near masker onset. The present study examined the effects of stimulus level and relative frequency region on this recovery. In all experiments, the masker was a broadband of noise and the signal was a 4-kHz sinusoid. The first experiment examined the effects of masker level on overshoot in order to choose two levels (one "low" and one "high") that produced similar amounts of overshoot; these levels were used in the remaining experiments. In the second experiment, the precursor was identical to the masker, and recovery functions were measured for both low and high masker and precursor levels. there was no consistent difference in the recovery functions between the two levels. In the third experiment, the precursor was divided into two bands (one below and one above 4 kHz); one was presented continuously while the other was gated as in experiment 2. The recovery was more complete when the band above 4 kHz was gated, although the recovery was usually less than that observed in experiment 2 when (effectively) both bands were gated. The results suggest that the frequency regions on both sides of the signal are important for the recovery of overshoot, but that the frequency region above the signal may be more important than the region below.

Some factors influencing comodulation masking release and across-channel masking.

The purpose of this study was to determine whether comodulation masking release (CMR) and across-channel masking (ACM) are by-products of a similar across-channel mechanism. This was addressed by examining how the two are affected by stimulus manipulations expected to influence their magnitude. Subjects were required to detect a 1000-Hz signal in the presence of a masker that consisted of a 1000-Hz (on-frequency) component alone or that component and up to six flanking components (500, 600, 700, 1300, 1400, and 1500 Hz). The on-frequency and flanking components typically were sinusoidally amplitude modulated at 10 Hz, although not necessarily in phase with one another. In experiment 1, the amount of CMR and ACM was highly influenced by whether the signal consisted of one or three 50-ms tone bursts; in fact, ACM was only observed when the signal was a train of three 50-ms tone bursts. In experiments 2 and 3, CMR tended to increase as the modulation depth or the number of flanking components increased, whereas ACM was relatively unaffected by these manipulations. In addition, ACM was observed under dichotic situations, whereas CMR was not. Taken together, the results suggest that ACM and CMR may be mediated by different mechanisms.

Effects of combining maskers in modulation detection interference.

The threshold for detecting 10-Hz amplitude modulation of a 2000-Hz carrier was measured in quiet, in the presence of an unmodulated masker, and in the presence of an amplitude-modulated masker. Two experiments were run; in each, the masker consisted of one or two sinusoidal carriers (chosen from among the frequencies of 800, 1600, 2400, and 3200 Hz). In experiment 1, the modulation rate of the masker ranged from 2 to 80 Hz. The "tuning" in the modulation domain was not affected much by the masker carrier frequency or the increase from one to two carriers. The amount of interference, however, was sometimes greater in the two-carrier condition, although this resulted primarily from the presence of the carriers and not from their modulation. In experiment 2, the modulation rate of each single-carrier masker ranged from 2 to 80 Hz (as in experiment 1), but for the two-carrier conditions, all possible combinations of two carriers (2400 and 3200 Hz) and three masker rates (5, 10, and 20 Hz) were evaluated. In general, the combination of two modulated carriers did not produce more interference than that produced by the more interfering carrier presented alone. Thus the results from both experiments provide little evidence for an additivity of modulation detection interference.