Okalianecopinata sp. nov. (Insecta, Coleoptera, Elmidae) from Gunung Mulu National Park in Sarawak (Malaysia).

Okalianecopinata sp. nov., from Sarawak, northwest Borneo, Malaysia, is described and illustrated along with an identification key. The standard barcoding fragment of the mitochondrial gene coding for cytochrome c oxidase subunit I (COI) was used together with morphological characters to delimit the taxonomic boundaries of the two known species, which live in shallow streams flowing through dense primary forests in limestone areas in Pahang (West Malaysia) and Sarawak (East Malaysia). The majority of all examined Okalia are flightless. Morphological distinguishing characters are the length of the granulated fifth elytral interval, the elytral and pronotal punctation, the aedeagal morphology, and the distal portion of the ovipositor.

Ancyronyx clisteri, a new spider riffle beetle species from Borneo, redescription of A. sarawacensis Jäch including a description of the larva and new distribution data for A. procerus Jäch using DNA barcodes (Coleoptera, Elmidae).

Ancyronyx clisteri sp. nov. (Coleoptera, Elmidae) a new spider riffle beetle discovered from northern Borneo (Brunei; Sabah and Sarawak, Malaysia) and the larva of Ancyronyx sarawacensis Jäch are described. Illustrations of the habitus and diagnostic characters of the new species and the similar and highly variable A. sarawacensis are presented. Differences to closely related species, based on DNA barcodes and morphological characters, are discussed. Association of the larva and the imago of A. sarawacensis, and the occurrence of Ancyronyx procerus Jäch in Peninsular Malaysia and Sabah are confirmed by using COI mtDNA sequences.

Ancyronyx lianlabangorum sp. nov., a new spider riffle beetle from Sarawak, and new distribution records for A. pulcherrimus Kodada, Jäch & Ciampor based on DNA barcodes (Coleoptera, Elmidae).

Ancyronyx lianlabangorum sp. nov. (Coleoptera, Elmidae), a new spider riffle beetle from the Kelabit Highlands (Sarawak, northern Borneo), is described. Illustrations of the habitus and diagnostic characters of the new species and the similar, polymorphic A. pulcherrimus Kodada et al. are presented. Differences to closely related species, based on COI nucleotide sequences and morphological characters, are discussed. Ancyronyx pulcherrimus is here recorded from Sarawak for the first time, based on DNA barcoding.

Revision of the species Grouvellinus Champion, 1923 (Coleoptera: Elmidae) with long median pronotal carina, including descriptions of four new species from China.

Four new species of Grouvellinus Champion, 1923 with very long median pronotal carina are described from China: G. hongkongensis sp. nov. (Guangdong, Guangxi, Hong Kong), G. longiusculus sp. nov. (Jiangxi), G. mediocarinatus sp. nov. (Fujian, Guangdong), and G. robustus sp. nov. (Anhui). These species are obviously closely related to G. bishopi Jäch, 1984 described from Malaysia (Selangor). The latter is here recorded for the first time from Kedah and Perak (Malaysia). Habitus photographs, detailed line drawings of the male genitalia, as well as a key to the males of the species of Grouvellinus with very long median pronotal carina are provided.

Stenelmis koreana Satô, 1978 (Coleoptera: Elmidae): confirmed as a wide-spread species by DNA-sequencing.

Stenelmis koreana Satô, 1978 (Coleoptera: Elmidae) is here recorded for the first time from Kyrgyzstan and Western Siberia. It was hitherto thought to be confined to Korea and the Russian Far East. The identification of a specimen from Kyrgyzstan was confirmed by DNA-sequencing after comparison with two sequences of S. koreana from Korea. The COI haplotype of the Kyrgyzstan specimen has very low sequence divergence (0.53 % or 0.0053 uncorrected p-distance) with respect to the sequences of the Korean specimens, which is within the standard intraspecific sequence divergence for COI in beetles.

Revision of the Chinese species of the genus Grouvellinus Champion, 1923 (Coleoptera: Elmidae). The G. acutus species group.

The Grouvellinus acutus species group is established. In China, it includes three species, Grouvellinus acutus sp. nov. (Fujian, Guangxi), G. denticulatus sp. nov. (Yunnan), and G. fujianensis sp. nov. (Fujian, Guangxi), which are described herein. A possible case of hybridization is reported. Habitus photographs and detailed line drawings of the genitalia are provided.

Hydraena Kugelann, 1794 (Coleoptera, Hydraenidae) from the Seychelles, Indian Ocean, with description of a new species.

Hydraena matyotisp. n. (Coleoptera, Hydraenidae) is described from the Seychelles, Indian Ocean. Hydraena mahensis Scott, 1913 is redescribed. The latter is here recorded from La Digue for the first time. A key to the species of the genus Hydraena Kugelann, 1794 of the Seychelles is presented.

Limnebius (Limnebius) agnieszkae sp. n. from the Crimean Peninsula (Coleoptera: Hydraenidae).

Limnebius (s.str.) agnieszkae sp. n. from the Crimean Peninsula (Ukraine) is described. It is a member of the Limnebius furcatus subgroup of the L. parvulus species group. The new species is very similar to L. stagnalis Guillebeau, 1890, from which it can be distinguished by the aedeagus only. Information about its habitat and a list of other water beetles found together with the new species are provided. Limnebius myrmidon Rey, 1883 is recorded for the first time from the Ukraine.

Electrolichas circumbalticus gen. et sp. nov. (Coleoptera: Byrrhoidea: Ptilodactylidae) from Baltic amber, the first anchytarsine toed-winged beetle described from Europe.

A new genus and species of Ptilodactylidae, Electrolichas circumbalticus gen. et sp. nov., the first fossil representative of the subfamily Anchytarsinae, is described from Baltic amber. This Eocene beetle can be distinguished from the extant genera of Anchytarsinae by the combination of the following characters: labrum transverse, terminal segment of maxillary palps cylindrical, antennae with non-articulated rami, basal pronotal margin only very finely crenulate, lacking a median denticle, anterior margin of scutellum not excised medially, elytra not striate, tarsi not pseudotetramerous, proportional lengths: 1.5: 1.2: 1.2: 1.1: 1.5, flashy pads on apex of first tarsomere as well as on tarsomeres 2-4. A list of the extant species of Anchytarsinae is provided. Both hitherto described fossil ptilodactylid genera are briefly discussed.

Annotated checklist of Dryopidae and Heteroceridae (Coleoptera: Byrrhoidea) of Iran.

The distribution of Dryopidae and Heteroceridae (Coleoptera: Byrrhoidea) of Iran is summarized. A total of ten species of Dryopidae in three genera (Dryops Olivier, Parahelichus Löbl & Smetana, and Praehelichus Löbl & Smetana), and 18 species of Heteroceridae in two genera (Augyles Schiödte and Heterocerus Fabricius) are listed. Augyles hispidulus (Kiesenwetter, 1843) (Heteroceridae) is recorded from Iran for the first time.

The riffle beetles (Coleoptera: Elmidae) of the Eocene Baltic amber: Heterelmis groehni sp. nov. and Heterlimnius samlandicus (Bollow, 1940) comb. nov.

The Eocene elmid beetles known from Baltic amber so far are reviewed, and their natural environment is discussed. Palaeoriohelmis samlandica Bollow, 1940 is re-described based on examination of the holotype. The genus Palaeoriohelmis Bollow, 1940 is synonymized with Heterlimnius Hinton, 1935, and Palaeoriohelmis samlandica is transferred to Heterlimnius (H. samlandicus comb. nov.). Heterelmis groehni sp. nov. is described and illustrated. Elmadulescens rugosus Peris, Maier & Sánchez-García, 2015 from Cretaceous Spanish amber is removed from Elmidae.

Fauna Europaea: Coleoptera 2 (excl. series Elateriformia, Scarabaeiformia, Staphyliniformia and superfamily Curculionoidea).

Fauna Europaea provides a public web-service with an index of scientific names (including synonyms) of all living European land and freshwater animals, their geographical distribution at country level (up to the Urals, excluding the Caucasus region), and some additional information. The Fauna Europaea project covers about 230,000 taxonomic names, including 130,000 accepted species and 14,000 accepted subspecies, which is much more than the originally projected number of 100,000 species. This represents a huge effort by more than 400 contributing specialists throughout Europe and is a unique (standard) reference suitable for many users in science, government, industry, nature conservation and education. Coleoptera represent a huge assemblage of holometabolous insects, including as a whole more than 200 recognized families and some 400,000 described species worldwide. Basic information is summarized on their biology, ecology, economic relevance, and estimated number of undescribed species worldwide. Little less than 30,000 species are listed from Europe. The Coleoptera 2 section of the Fauna Europaea database (Archostemata, Myxophaga, Adephaga and Polyphaga excl. the series Elateriformia, Scarabaeiformia, Staphyliniformia and the superfamily Curculionoidea) encompasses 80 families (according to the previously accepted family-level systematic framework) and approximately 13,000 species. Tabulations included a complete list of the families dealt with, the number of species in each, the names of all involved specialists, and, when possible, an estimate of the gaps in terms of total number of species at an European level. A list of some recent useful references is appended. Most families included in the Coleoptera 2 Section have been updated in the most recent release of the Fauna Europaea index, or are ready to be updated as soon as the FaEu data management environment completes its migration from Zoological Museum Amsterdam to Berlin Museum für Naturkunde.

Anisomeriini diving beetles-an Atlantic-Pacific Island disjunction on Tristan da Cunha and Robinson Crusoe Island, Juan Fernández?

Anisomeriini diving beetles contain only two enigmatic species, representing a remarkable disjunction between the Pacific Juan Fernández Islands (Anisomeria bistriata) and the South Atlantic Tristan da Cunha Archipelago (Senilites tristanicola). They belong to the Colymbetinae, which contain 140 species worldwide. Here we aim to reconstruct the evolutionary history of the Anisomerinii and use > 9000 bp DNA sequence data from 13 fragments of 12 loci for a comprehensive sampling of Colymbetinae species. Analyses under different optimization criteria converge on very similar topologies, and show unambiguously that Anisomeria bistriata and Senilites tristanicola belong to the Neotropical Rhantus signatus species group, a comparatively recent clade within Colymbetinae. Anisomeriini therefore are synonomized with Colymbetini and both species are transferred to Rhantus accordingly, resulting in secondary homonymy of Rhantus bistriatus (Brullé, 1835) with Rhantus bistriatus (Bergsträsser, 1778). We propose the replacement name Rhantus selkirki Jäch, Balke & Michat nom. nov. for the Juan Fernández species. Presence of these species on remote islands is therefore not relictary, but the result of more recent range expansions out of mainland South America. Finally, we suggest that Carabdytini should be synonymized with Colymbetini. Our study underpins the Hennigian principle that a natural classification can be derived only from the search for shared apomorphies between species, not from differences.

Ancyronyx reticulatus and A. pulcherrimus, two new riffle beetle species from Borneo, and discussion about elmid plastron structures (Coleoptera: Elmidae).

Two new species of Ancyronyx Erichson, 1847 (Coleoptera: Elmidae) are described from Borneo: A. pulcherrimus (Brunei) and A. reticulatus (Sabah). Habitus views, illustrations of important characters as well as plastron structures of Ancyronyx reticulatus are presented and discussed.

Two new species of Ochthebius Leach (Coleoptera: Hydraenidae) from Turkey.

Two new species of Ochthebius Leach (Coleoptera: Hydraenidae) are described from Turkey: O. (s.str.) halophilus (Kirikkale Province) and O. (Asiobates) arator (Kütahya and Usak Provinces). The former is the first member of the Ochthebius (s.str.) notabilis species group from Turkey, and the latter belongs to the O. (Asiobates) bicolon group. Both of the new species are characterized by a remarkable aedeagus and are very probably endemic to Turkey.

Ochthebius hivae (Coleoptera: Hydraenidae) from western Iran, a new species of the O. metallescens group with remarkable aedeagal variability.

Ochthebius hivae is described from western Iran (Ilam and Khuzestan Provinces). It belongs to the O. metallescens Rosenhauer, 1847 species group. The aedeagal distal lobe of this species is remarkably variable.