Review of the leafhopper genus Drabescus Stl (Hemiptera: Cicadellidae: Deltocephalinae) in the Indian subcontinent with description of one new species.

The leafhopper genus Drabescus Stl is reviewed for the Indian subcontinent with one new species described, D. austroindicus sp. nov. (India: Tamil Nadu). A checklist and key to the species from the Indian subcontinent are given.

Four new species of Mileewini leafhoppers (Hemiptera: Cicadellidae: Mileewinae) from China, with a checklist to Chinese species.

Four new species of Mileewini, Mileewa clavata He Yang, sp. nov., Mileewa rubricosta He Yang, sp. nov., Mileewa quinquemaculata He Yang, sp. nov. and Ujna liboensis He Yang, sp. nov., are described and illustrated. A checklist of all Chinese Mileewini is provided.

Review of the leafhopper genus Protensus/ Zhang amp; Dai (Hemiptera, Cicadellidae, Deltocephalinae) with description of a new species.

Species of the leafhopper genus Protensus Zhang Dai, 2001 (Deltocephalinae: Athysanini) from Japan and China are reviewed, and a new species Protensus lii sp. nov. is described and illustrated from China. Protensus nigrifrons Li Xing, 2011 is treated as incertae sedis in the tribe Mukariini (Deltocephalinae). A checklist and key to species of Protensus is provided.

Leafhopper subfamily Ledrinae of the Indian subcontinent: 1. Description of Yelahanka gen. nov. (Hemiptera: Cicadellidae) with seven new species and new combinations.

One new genus of the tribe Ledrini, Yelahanka gen. nov. is described with Petalocephala granulosa Distant as its type species. The genus is characterised by the flexing of the forewing laterad of a strongly carinate or keeled outer claval vein and the claval veins fused in the distal two thirds. Seven new species, Yelahanka canaraica sp. nov. (India: Karnataka), Y. kodaiensis sp. nov. (India: Tamil Nadu), Y. montana sp. nov. (India: Tamil Nadu), Y. nepalica sp. nov. (Nepal), Y. sikkimensis sp. nov. (India: Sikkim), Y. shillongensis sp. nov. (India: Meghalaya) and Y. trifida sp. nov. (India: Himachal Pradesh, Arunachal Pradesh) are described and illustrated. The following new combinations are proposed: Yelahanka bainbriggei (Distant) comb. nov., Y. granulosa (Distant) comb. nov., Y. punctata (Walker) comb. nov. and Y. tabulata (Distant) comb. nov.; originally punctata was placed in the genus Ledra Fabricius and remaining species were placed in Petalocephala Stål. Petalocephala bainbriggei Distant 1916 is treated as a junior synonym of Y. granulosa (Distant 1910) syn. nov. and P. tabulata Distant 1908 is treated as a junior synonym of Yelahanka punctata (Walker 1851) syn. nov. Y. granulosa is newly recorded from Africa (Tanzania and Kenya) and Malaya. Relationships of the new genus with other genera of Ledrini are discussed and a key to species of the genus is also provided.

A new species of the Oriental leafhopper genus Kutara Distant (Hemiptera: Cicadellidae: Deltocephalinae) from Vietnam.

The Oriental leafhopper genus Kutara Distant, 1908 (Cicadellidae: Deltocephalinae: Drabescini: Paraboloponina) is newly recorded from Vietnam with description of a new species K. trispinosa Xu Zhang sp. nov.. An updated checklist, with distribution and a key to males of Kutara, are also provided. The type specimens of the new species are deposited in the Muséum National d'Histoire Naturelle, Paris, France (MNHN) and the Illinois Natural History Survey, University of Illinois, Champaign, Illinois, USA (INHS).

Leafhopper tribe Stegelytrini (Hemiptera: Cicadellidae: Deltocephalinae) of the Indian subcontinent, with a note on Aeternus hieroglyphicus Distant (Cicadellidae: Athysanini).

A checklist of leafhoppers of the tribe Stegelytrini from the Indian subcontinent and a key to genera are given.  Two new species, Pachymetopius anamikus sp. nov. (India: Meghalaya) and Yaontogonia unipuncta sp. nov. (India: Arunachal Pradesh) are described and illustrated. Genus Aeternus Distant with its type species A. hieroglyphicus Distant is redescribed based on the male genitalia and its tribal placement is discussed. Habitus images of types of species except Aculescutellaris hirtellus Zhang, Wei Webb are given.

A checklist and key to the idiocerine leafhoppers (Hemiptera: Cicadellidae) of Hainan Island, with description of a new genus and new species.

The leafhopper subfamily Idiocerinae from Hainan Island (China) is reviewed. Eight genera and 10 species are recognized including one new genus and one new species, Longistyla gen. nov. and Longistyla viraktamathi sp. nov., based on specimens from Jianfengling Nature Reserve. A checklist and keys to genera and species are included.

On the identity and distribution of the Old World grass feeding leafhopper species Soractellus nigrominutus Evans (Hemiptera: Cicadellidae: Deltocephalinae: Paralimnini).

The identity of the Old World grass feeding leafhopper species Soractellus nigrominutus Evans, 1966 is clarified and its distribution revised. Based on examination of material, from throughout its range, the following species are placed as new subjective junior synonyms of S. nigrominutus: S. jianfengensis Xing Li, 2014, syn. nov., and S. lalianensis Naveed Zhang, 2018, syn. nov. The following new country records for S. nigrominutus are recorded: Thailand, Nigeria, Kenya, Uganda, Senegal, Ethiopia and Saudi Arabia.

Revision of the Ulopinae leafhoppers (Hemiptera: Cicadellidae) of the Indian subcontinent, I. Ulopini genera: Daimachus, Radhades and Ulopsina.

The Ulopinae leafhopper genera of the tribe Ulopini from the Indian subcontinent (Daimachus Distant and Radhades Distant) are redescribed. Four new species of Daimachus are described: D. matheranensis sp. nov. (India: Maharashtra), D. robustus sp. nov. (India: Kerala), D. sirsiensis sp. nov. (India: Karnataka) and D. sudindicus sp. nov. (India: Karnataka, Kerala). Ulopa brunnea Pruthi 1930 is treated as a junior synonym of Daimachus exemplificatus Distant 1916 based on a study of the male genitalia of both species. The latter species, together with Radhades crassus Distant and Ulopsina Dai, Viraktamath Zhang are redescribed and a new subgenus of Ulopsina, Indoulopa subgen. nov. is described from Sub-Himalayan region with Ulopsina (Indoulopa) himalayana sp. nov. as the type species; the genus is reported for the first time from the subcontinent. Key to genera and species dealt with are also provided, all taxa are illustrated and lectotypes are designated for Daimachus exemplificatus, Ulopa brunnea and Radhades crassus.

Two new genera of Pacific Paraboloponina leafhoppers (Hemiptera: Cicadellidae: Drabescini) with description of three new species.

Two new genera, Malaysiapona gen. nov. and Chandrapona gen. nov. of Paraboloponina leafhoppers (Cicadellidae: Deltocephalinae: Drabescini) with their unusally developed aedeagal base are described from the Pacific (Malaysia). The following new species are also described with the new genera: Malaysiapona brevipenis sp. nov. from Sabah, M. filamenta sp. nov., from Sarawak and Chandrapona vespertilis sp. nov. from Sabah.

Revision of the bamboo leafhopper tribe Mukariini (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent with description of new genera and species.

Leafhopper genera and species of the tribe Mukariini from the Indian subcontinent are revised. Nine genera and 22 species including two new genera, one new subgenus and 12 new species are dealt with. The new taxa described are Aalinga gen. nov. with its type species Aalinga brunoflava sp. nov. (India: Andaman Islands), Buloria indica sp. nov. (India: Karnataka). Buloria zeylanica sp. nov. (Sri Lanka), Flatfronta bella sp. nov. (India: Karnataka; Bangladesh), Mohunia bifurcata sp. nov. (Myanmar), Mukaria omani sp. nov. (India: Gujarat, Karnataka, Madhya Pradesh), Mukaria vakra sp. nov. (India: Karnataka), Mukariella gen. nov. with its type species Mukariella daii sp. nov. (India: Manipur), Myittana (Benglebra) cornuta sp. nov. (India: Karnataka), Myittana (Myittana) distincta sp. nov. (India: Karnataka), Myittana (Savasa) subgen. nov. with its type species Myittana (Savasa) constricta sp. nov. (India: Andhra Pradesh, Gujarat, Haryana, Himachal Pradesh, Karnataka, Madhya Pradesh, Punjab, Tamil Nadu, Uttarakhand) and Scaphotettix arcuatus sp. nov. (India: West Bengal, Meghalaya, Mizoram). Genera Buloria Distant (new placement), Crispina Distant (new placement) and Myittana Distant (new placement) are placed in the tribe Mukariini. Genus Mohunia is redefined based on the study of its type species. Benglebra Mahmood Ahmed 1969 is synonymised with Myittana Distant 1908 and considered as its subgenus. Myittana (Benglebra) alami (Mahmood Ahmed) comb. nov., Myittana (Savasa) bipunctata (Mahmood Ahmed) comb. nov.. Myittana (Benglebra) introspina (Chen Yang 2007) comb. nov. and Mukariella bambusana (Li Chen) comb. nov. are proposed; the first two species were earlier placed in the genus Benglebra, the third species in the genus Mohunia and the fourth in the genus Mukaria. Genera Flatfronta Chen Li and Myittana are new records for India and Scaphotettix striata Dai Zhang is a new record for the Indian subcontinent and Sri Lanka. All taxa dealt with are described and illustrated and keys for genera and their species are also given.

A new leafhopper genus of subtribe Paraboloponina (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini), with description of two new species from Papua New Guinea.

A new leafhopper genus Papuakutara gen. nov. is described from Papua New Guinea, with two new species, P. ficus sp. nov. and P. robustipenis sp. nov. Kutara lucidicosta (Walker, 1870) is transferred to Papuakutara comb. nov. A checklist of the genera in the subtribe Paraboloponina from Papua New Guinea and West Papua and a key for their separation are provided.

Revision of the penthimiine leafhopper genus Tambila Distant (Hemiptera: Cicadellidae: Deltocephalinae) of the Indian subcontinent.

The genus Tambila Distant of the deltocephaline tribe Penthimiini of the Indian Subcontinent is revised. Tambila opulenta Distant is placed as a junior synonym of Tambila greeni Distant and Tambila gravelyi Distant as a junior synonym of Tambila majuscula (Distant). All valid taxa are redescribed, illustrated and a key to the eight known species from the subcontinent is included. A new record is given for T. gravely from Mauritius (on mango).

Description of two new idiocerine leafhoppers of the genus Chinaocerus (Hemiptera: Cicadellidae) from southwestern China.

The leafhopper genus Chinaocerus Zhang Li (Cicadellidae: Idiocerinae) currently includes three species, C. bispinatus Zhang Li (type species), C. kangdingensis Zhang Li and C. shii Zhang Li from southwestern China. Two new species, C. tubulatus Zhang Webb sp. nov. and C. sexspinosus Zhang Webb sp. nov. from Yunnan Province, China, are here described and a checklist and key to species are provided.

Review of the leafhopper genus Penthimia Germar (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent with description of seven new species.

Species of the leafhopper genus Penthimia Germar known from the Indian subcontinent are reviewed based on the examination of type specimens. Seven new species of the genus, Penthimia curvata sp. nov. (Karnataka: Bandipur), P. meghalayensis sp. nov. (Meghalaya: Nangpoh), P. neoattenuata sp. nov. (India: Tamil Nadu), P. ribhoi sp. nov. (India: Meghalaya), P. sahyadrica sp. nov. (Karnataka: Dharmasthala, Agumbe; Kerala: Thekkady), P. spiculata sp. nov. (Karnataka: Nagarahole) and P. tumida sp. nov. (Tamil Nadu: Ootacamund; Kerala: Munnar) are described. The following nomenclatorial changes are proposed: Penthimia alba Zahniser, McKamey Dmitriev, 2012 (replacement name for P. thoracica Distant, 1918, nec Panzer, 1799), syn. nov. of P. quadrinotata Distant, 1918; Neodartus scutellatus Distant, 1908 syn. nov. of Penthimia ereba Distant 1908; P. nilgiriensis Distant, 1918 syn. nov. of P. montana Distant, 1918; P. scutellata (Distant) comb. nov. (from genus Neodartus); a lectotype is designated for P. maculosa Distant, stat. revived, thereby removing its synonymy with P. scapularis Distant. The following other lectotypes are designated: P. attenuata Distant, P. subniger Distant, P. scapularis Distant, P. distanti Baker, P. ereba Distant, N. scutellatus Distant, P. fraterna Distant, P. funebris Distant, P. juno Distant, P. maculosa Distant, P. montana Distant, P. noctua Distant, P. quadrinotata Distant, P. alba Zahniser, McKamey Dmitriev. Examination of types of Penthimia rufopunctata Motschulsky revealed that it belongs to Penthimia and hence it is transferred back to that genus from Neodartus, revised placement. The following species previously included in the genus Penthimia are transferred to the genera Tambila Distant and Vulturnus Kirkaldy: Tambila badia (Distant) comb. nov., T. majuscula (Distant) comb. nov., T. vittatifrons (Distant) comb. nov., T. variabilis (Distant) comb. nov. and Vulturnus flavocapitata (Distant) comb. nov. Three species are treated in a new Penthimia compacta Walker complex, i.e., Penthimia compacta Walker 1851, Penthimia subniger Distant 1908 and Penthimia scapularis Distant 1908. All taxa are described and a key to Penthimiini genera found in the subcontinent and also a key to species of Penthimia are included.

Revision of the evacanthine leafhoppers (Hemiptera: Cicadellidae: Evacanthinae) of the Indian subcontinent.

The leafhoppers of the tribe Evacanthini of the Indian subcontinent (Pakistan, India, Nepal, Bhutan, Bangladesh, Myanmar and Sri Lanka) are reviewed. The genus Apphia Distant 1918, is reinstated as a valid genus from synonymy with Onukia Matsumura stat. rev. and redefined based on the male genitalia characters. The genus Angustuma Xing Li 2013, (new name for Angustella Li 1986, not Waagen 1907) is considered a junior synonym of Apphia. The genera Carinata Li Wang, Concavocorona Wang Zhang, Risefronta Li Wang, Shortcrowna Li Li, Striatanus Li Wang and Taperus Li and Wang along with Bundera bambusana Yang, Chen Li, Carinata midfasciana Wang Li, Onukia albiclypeus Li Wang, O. flavocapitata Li Wang and Risefronta albicincta Li Wang are here recorded for the first time from the Sub-Himalayan region of the Indian subcontinent. Keys to genera and included species are given. The new taxa recognized are Apphia flava sp. nov. (India: Meghalaya), Bundera acutivertex sp. nov. (India: Mizoram), Concavocorana kambaiti sp. nov. (Myanmar; Kambaiti), Evacanthus albipennis sp. nov. (Nepal), E. convolutus sp. nov. (India: Himachal Pradesh), E. distinctus sp. nov. (India: Himachal Pradesh), E. manaliensis sp. nov. (India: Himachal Pradesh), E. yeshwanthi sp. nov. (India: Meghalya), Onukia echina sp. nov. (India: Manipur), Onukindia gen. nov. (type species : Platyretus connexus Distant), Onukindia aruna sp. nov. (India: Arunachal Pradesh), O. compressa sp. nov. (India: Arunachal Pradesh, West Bengal), Shortcrowna rubrostriata sp. nov. (Myanmar: Kambaiti), Striatanus delta sp. nov. (India: West Bengal; Mizoram) and Taperus indicus sp. nov. (India: Arunachal Pradesh). The following new combinations (genera of original combination in parenthesis): Apphia albida Li Li) comb. nov., A. albonotata (Yang Zhang) comb. nov., A. basinigra (Wang Zhang) comb. nov., A. flavimacula (Wang Zhang) comb. nov., A. flavinigra (Wang Zhang) comb. nov., A. jinghongensis (LI Li) comb. nov., A. leucostriata (Li Wang) comb. nov., A. longipyga (Li Wang) comb. nov., A. nigricarina (Li) comb. nov., A. nigricauda (Cheng, Yang Li) comb. nov., A. nigrimargina (Li Li) comb. nov., A. nigrinota (Yang Zhang) comb. nov., A. pallida (Cai He) comb. nov., A. panxianensis (Li Li) comb. nov., A. rudorsuma (Xing Li) comb. nov. and A. rufipenna (Li Wang) comb. nov. (all from Angustuma), Apphia assamensis (Ramakrishnan) comb. nov. (Dussana), Apphia himalayana (Distant) comb. nov. (Platyretus) and Onukindia connexa (Distant) comb. nov. (Platyretus) are made. Evacanthus repexus Distant and Vangama steneosaura Distant appear to be species complexes and need to be further investigated. Lectotypes are designated for: Apphia burmanica Distant, Apphia himalayana (Distant), Bundera venata Distant, Cunedda phaeops Distant, Evacanthus bellus (Distant), Evacanthus militaris (Distant), Evacanthus repexus Distant, Mainda praeculta Distant, Onukindia connexa (Distant) and Vangama steneosaura Distant.

Punctulini (Hemiptera: Cicadellidae: Deltocephalinae), a new leafhopper tribe from the Oriental region and Pacific Islands.

A new tribe of the leafhopper subfamily Deltocephalinae, Punctulini tribe nov., from China, India, Thailand, and Fiji is described and includes three new genera and five new species: Punctulus gen. nov. includes P. tumidifrons sp. nov. from China, P. manipurensis from India, and P. lobatus sp. nov. from Thailand; Hirsutula gen. nov. includes H. rubrifrons sp. nov. from Thailand; and Taveunius gen. nov. includes T. megapunctatus sp. nov., from Fiji. A key to the genera of Punctulini is provided. Observations on the shared morphological characters of Punctulini with the superficially similar tribe Magnentiini, and with Vartini to which it is found to be related based on phylogenetic analyses of morphological and molecular data including newly available sequences and analyses, are made and discussed. Afrovarta Zahniser 2013 is removed from Vartini and placed in Athysanini, placement nov.

Review of the leafhopper tribe Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) from China with description of four new species.

The leafhopper tribe Stenometopiini is revised from China. Two genera are included: Hodoedocus Jacobi with two species and Stirellus Osborn & Ball with thirteen species of which four are new: S. daii sp. nov., S. multipunctatus sp. nov., S. neospeciosus sp. nov., and S. translucidus sp. nov. S. yeongnamensis (Kwon & Lee) is newly recorded for China. Five new synonyms are proposed: Viridomarus brevialatus Xing et al. and Viridomarus laticellus Xing et al. as junior synonyms of S. capitatus (Distant); Aconura bipunctella Matsumura and Paivanana centristriata Dai & Li as junior synonyms of S. indrus (Distant); Aconura rubrofasciata Matsumura as junior synonym of S. rubrolineatus (Distant). Two new combinations are erected and one combination is reinstated (see checklist). Species are described and illustrated together with keys provided for their separation. Lectotypes are designated for Typhlocyba indra Distant and Arya rubrolineata Distant.

A new genus and species of the leafhopper subtribe Paraboloponina from China (Hemiptera, Cicadellidae).

A new leafhopper genus and species Forficus maculatus Qu gen. nov., sp. nov. of the subtribe Paraboloponina (Deltocephalinae, Drabescini) are described from China. A checklist to the Paraboloponina from China is given and a key to genera is provided.

Review of the idiocerine leafhoppers of Pakistan (Hemiptera, Cicadellidae) with a description of a new species.

The Idiocerinae of Pakistan are reviewed and a new species, Tasnimocerus sindhensis sp. nov. (Pakistan: Tandojam), is described and illustrated from Pakistan. Two new junior synonyms of Idioscopus nitidulus (Walker) are recognized: Idioscopus karachiensis Ahmed, Naheed & Ahmed syn. nov. and I. freytagi Ahmed, Naheed & Ahmed syn. nov. Idioscopus nagpurensis (Pruthi) is newly recorded from Pakistan. A checklist of Idiocerinae from Pakistan is also provided together with a key to genera and species.