Behavioural Indicators of Intra- and Inter-Specific Competition: Sheep Co-Grazing with Guanaco in the Patagonian Steppe.

In extensive livestock production, high densities may inhibit regulation processes, maintaining high levels of intraspecific competition over time. During competition, individuals typically modify their behaviours, particularly feeding and bite rates, which can therefore be used as indicators of competition. Over eight consecutive seasons, we investigated if variation in herd density, food availability, and the presence of a potential competitor, the guanaco (Lama guanicoe), was related with behavioural changes in domestic sheep in Chilean Patagonia. Focal sampling, instantaneous scan sampling, measures of bite and movement rates were used to quantify behavioural changes in domestic sheep. We found that food availability increased time spent feeding, while herd density was associated with an increase in vigilant behaviour and a decrease in bite rate, but only when food availability was low. Guanaco presence appeared to have no impact on sheep behaviour. Our results suggest that the observed behavioural changes in domestic sheep are more likely due to intraspecific competition rather than interspecific competition. Consideration of intraspecific competition where guanaco and sheep co-graze on pastures could allow management strategies to focus on herd density, according to rangeland carrying capacity.

Ingestive behavior of Girolando heifers in integrated crop, livestock (ICL), and forestry (ICLF) systems / [Comportamento ingestivo de novilhas Girolando mantidas em sistemas de integração lavoura-pecuária (ILP) e lavoura-pecuária-floresta (ILPF]

Aiming to compare integrated crop, livestock (ICL) and forest (ICLF) systems in relation to ingestive behavior of dairy heifers, a 2 × 2 crossover trial was conducted with two periods and two systems. Eight Girolando (3/4holstein × 1/4 Gyr) heifers were evaluated in pasture of Xaraés palisade grass (Urochloa brizantha 'Xaraes' syn Brachiaria brizantha) managed with intermittent stocking. The ICLF systemhad eucalyptus trees planted in tiers with 65% crown cover. Ingestive behavior parameters were obtained using acoustic data collected with mP3 recorders for 48hours. Statistical analysis was performed using the mixed procedure of SAS, and means were compared using the Tukey-Kramer test at 5% significance. The dry mater intake (DMI), number of daily grazing sessions, and the duration of the grazing sessions did not differ (P<0.05) between systems. A higher (P<0.05) bite rate and total daily bites and lower (P<0.05) bite mass was observed in the ICLF system in relation to ICL. Heifers in shaded pasture change their ingestive behavior in comparison with those in full-sun pasture.(AU)

Com o objetivo de comparar sistemas de integração lavoura-pecuária (ILP) e lavoura-pecuária-floresta (ILPF) em relação ao comportamento ingestivo de novilhas leiteiras, conduziu-se um ensaio em delineamento crossover 2 × 2, com dois períodos e dois sistemas. Oito novilhas Girolando foram mantidas em pastagens de capim-xaraés (Urochloa brizantha 'Xaraes' syn Brachiaria brizantha) manejadas com lotação intermitente. No sistema ILPF, havia renques de eucalipto com 65% de cobertura de copa. O comportamento ingestivo foi avaliado pela análise de áudios coletados com gravadores de Mp3, durante 48 horas. As análises estatísticas foram realizadas utilizando-se Proc mixed SAS. As médias foram comparadas pelo teste Tukey-Kramer, ao nível de 5% de significância. A ingestão de matéria seca (IMS), o número de sessões de pastejo por dia e a duração das sessões de pastejo não diferiram (P<0,05) entre os sistemas. As maiores (P<0,05) médias de taxa de bocado e de número de bocados por dia e a menor (P<0,05) média de massa de bocado foram observadas no ILPF quando comparado ao ILP. Novilhas em pastagem sombreada alteram seu comportamento ingestivo quando comparadas àquelas que são mantidas em pastagem a pleno sol.(AU)

Sward structure and ingestive behavior of cows in tropical pastures managed under different forage allowances / Estrutura do dossel forrageiro e comportamento ingestivo de vacas em pastos tropicais manejados sob diferentes ofertas de forragem

This study evaluated how changing the structure of Brachiaria brizantha cv. Marandu pasture under different forage allowances (FA) of 4, 7, 10 and 13kg DM/100kg BW (body weight) affected animal behavior over a two-year evaluation period. The experiment was conducted as a randomized block design with three replications (paddock). Sward height, total forage, and stem mass were lower for pastures managed with lower FA. Lower leaf mass was observed for lower FA in the second year. In hand-plucked samples, leaf and stem percentages remained unchanged in the morning but leaf percentage increased while stems decreased in the afternoon. Permanence time decreased linearly with increasing FA. In turn, a quadratic effect was observed for displacement rate. The bite rate was similar among different FAs. The results demonstrate that FA varying between 7 and 10kg DM/100kg BW are more suitable to balance the amounts of leaves and stems in the sward. The studied FA levels do not limit forage availability. Permanence time decreases and displacement rate increases as FA increases.(AU)

O objetivo foi avaliar como mudanças na estrutura de pastos de Brachiaria brizantha cv. Marandu, sob diferentes ofertas de forragem (OF) de 4, 7, 10 e 13kg MS/100kg PC, afetam o comportamento animal durante um período de avaliação de dois anos. O delineamento utilizado foi em blocos ao acaso, com três repetições (piquetes). A altura do dossel, a massa de forragem total e de colmos foram menores em pastos manejados com menores OF. A menor massa de folhas foi observada na menor OF no segundo ano. Nas amostras de pastejo simulado, as porcentagens de folhas e colmos não diferiram pela manhã, mas a porcentagem de folhas aumentou, enquanto a de colmos diminuiu à tarde. O tempo de permanência dos animais diminuiu linearmente com o aumento das OF. Efeito quadrático foi observado para taxa de deslocamento. A taxa de bocados foi similar entre as OF. Os resultados demonstram que variar as OF entre 7 e 10kg MS/100kg PC é mais adequado para balancear as quantidades de folhas e colmos no dossel forrageiro. Os níveis de OF estudados não limitam a forragem disponível. O tempo de permanência diminuí e a taxa de deslocamento aumenta com o aumento das OF.(AU)

Sward structure and ingestive behavior of cows in tropical pastures managed under different forage allowances / Estrutura do dossel forrageiro e comportamento ingestivo de vacas em pastos tropicais manejados sob diferentes ofertas de forragem

This study evaluated how changing the structure of Brachiaria brizantha cv. Marandu pasture under different forage allowances (FA) of 4, 7, 10 and 13kg DM/100kg BW (body weight) affected animal behavior over a two-year evaluation period. The experiment was conducted as a randomized block design with three replications (paddock). Sward height, total forage, and stem mass were lower for pastures managed with lower FA. Lower leaf mass was observed for lower FA in the second year. In hand-plucked samples, leaf and stem percentages remained unchanged in the morning but leaf percentage increased while stems decreased in the afternoon. Permanence time decreased linearly with increasing FA. In turn, a quadratic effect was observed for displacement rate. The bite rate was similar among different FAs. The results demonstrate that FA varying between 7 and 10kg DM/100kg BW are more suitable to balance the amounts of leaves and stems in the sward. The studied FA levels do not limit forage availability. Permanence time decreases and displacement rate increases as FA increases.(AU)

O objetivo foi avaliar como mudanças na estrutura de pastos de Brachiaria brizantha cv. Marandu, sob diferentes ofertas de forragem (OF) de 4, 7, 10 e 13kg MS/100kg PC, afetam o comportamento animal durante um período de avaliação de dois anos. O delineamento utilizado foi em blocos ao acaso, com três repetições (piquetes). A altura do dossel, a massa de forragem total e de colmos foram menores em pastos manejados com menores OF. A menor massa de folhas foi observada na menor OF no segundo ano. Nas amostras de pastejo simulado, as porcentagens de folhas e colmos não diferiram pela manhã, mas a porcentagem de folhas aumentou, enquanto a de colmos diminuiu à tarde. O tempo de permanência dos animais diminuiu linearmente com o aumento das OF. Efeito quadrático foi observado para taxa de deslocamento. A taxa de bocados foi similar entre as OF. Os resultados demonstram que variar as OF entre 7 e 10kg MS/100kg PC é mais adequado para balancear as quantidades de folhas e colmos no dossel forrageiro. Os níveis de OF estudados não limitam a forragem disponível. O tempo de permanência diminuí e a taxa de deslocamento aumenta com o aumento das OF.(AU)

Performance and grazing behavior of growing goats supplemented with palm tree fruit

This study aimed to evaluate the performance and ingestive behavior of growing goats grazing on Tanzania guinea grass and fed diets containing 40% carnauba or tucum fruits. Twenty-one male castrated goats were distributed into three groups, one exclusively on pasture and the other two on pasture and fed diet supplemented at the level of 1.5% body weight (BW) in a completely randomized design. The intake of the supplements was obtained by difference between the amount supplied and the leftovers, with weighing performed every seven days, while pasture intake was determined using titanium dioxide (TiO2) as external indicator. Ingestive behavior was evaluated for three days. The supplement containing carnauba fruit resulted in a greater intake of neutral detergent fiber (0.137 kg NDF/day), with a reduction of 8.61% in the pasture dry matter (DM) intake of goats. Associated with the intake of pasture nutrients, the tucum fruit diet met the protein (0.103 kg CP/day) and energetic (0.547 kg TDN/day) requirements of goats with intake set at 0.124 kg CP/day and 0.572 kg TDN/day, with higher weight gain (0.111 kg/day) and larger loin eye area (12.76 cm2). The supplementation with fruits influenced the grazing behavior of goats, increasing the idle time by 1 h in relation to animals not supplemented. The supplementation of growing goats grazing on Tanzania guinea grass pasture with a diet containing 40% tucum fruit, in the proportion of 1.5% BW, did not meet the nutritional requirements for gain of 150 g/day; however, it met requirements for maintenance and average gain of 111 g/day. Energy supplementation reduces the grazing time of goats; thus, it is necessary to consider the level and formulation of supplements, with the possibility of increasing the stocking rate and productivity per unit area.(AU)

Quantifying seasonal and diel variation in Anopheline and Culex human biting rates in Southern Ecuador.

BACKGROUND: Quantifying mosquito biting rates for specific locations enables estimation of mosquito-borne disease risk, and can inform intervention efforts. Measuring biting itself is fraught with ethical concerns, so the landing rate of mosquitoes on humans is often used as a proxy measure. Southern coastal Ecuador was historically endemic for malaria (Plasmodium falciparum and Plasmodium vivax), although successful control efforts in the 2000s eliminated autochthonous transmission (since 2011). This study presents an analysis of data collected during the elimination period. METHODS: Human landing catch (HLC) data for three mosquito taxa: two malaria vectors, Anopheles albimanus and Anopheles punctimacula, and grouped Culex spp. were examined for this study. These data were collected by the National Vector Control Service of the Ministry of Health over a 5-year time span (2007-2012) in five cities in southern coastal Ecuador, at multiple households, in all months of the year, during dusk-dawn (18:00-6:00) hours, often at both indoor and outdoor locations. Hurdle models were used to determine if biting activity was fundamentally different for the three taxa, and to identify spatial and temporal factors influencing bite rate. Due to the many different approaches to studying and quantifying bite rates in the literature, a glossary of terms was created, to facilitate comparative studies in the future. RESULTS: Biting trends varied significantly with species and time. All taxa exhibited exophagic feeding behavior, and outdoor locations increased both the odds and incidence of bites across taxa. Anopheles albimanus was most frequently observed biting, with an average of 4.7 bites/h. The highest and lowest respective months for significant biting activity were March and July for An. albimanus, July and August for An. punctimacula, and February and July for Culex spp. CONCLUSIONS: Fine-scale differences in endophagy and exophagy, and temporal differences among months and hours exist in biting patterns among mosquito taxa in southern coastal Ecuador. This analysis provides detailed information for targeting vector control activities, and household level vector prevention strategies. These data were collected as part of routine vector surveillance conducted by the Ministry of Health, and such data have not been collected since. Reinstating such surveillance measures would provide important information to aid in preventing malaria re-emergence.

Mechanisms and implications of a type IV functional response for short-term intake rate of dry matter in large mammalian herbivores.

The functional response (i.e. the relationship between consumers' intake rate and resource density) is central in plant-herbivore interactions. Its shape and the biological processes leading to it have significant implications for both foraging theory and ecology of grazing systems. A type IV functional response (i.e. dome-shaped relationship) of short-term intake rate of dry matter (intake while grazing) has rarely been reported for large herbivores and the conditions that can lead to it are poorly understood. We report a type IV functional response observed in heifers grazing monocultures of Cynodon sp. and Avena strigosa. The mechanisms and consequences of this type of functional response for grazed system dynamics are discussed. Intake rate was higher at intermediate than at short or tall sward heights in both grass species. The type IV functional response resulted from changes in bite mass instead of a longer time needed to encounter and process bites. Thus, the decrease of intake rate of dry matter in tall swards is not explained by a shift from process 3 (potential bites are concentrated and apparent) to process 2 (potential bites are apparent but dispersed, Spalinger & Hobbs 1992). Bite mass was smaller in tall than in intermediate swards due to a reduction of bite volume possibly caused by the greater proportion of stem and sheath acting as a physical barrier to bite formation. It is generally accepted that potential bites are abundant and apparent in most grassland and meadow systems, as they were in the present experiments. Therefore, a type IV response of intake rate not directly related to digestive constraints may determine the dynamics of intake and defoliation under a much larger set of conditions than previously thought. These results have implications for foraging theory and stability of grazing systems. For example, if animals prefer patches of intermediate stature that yield the highest intake rate, grazing should lead to the widely observed bimodal distribution of plant mass per unit area, even when tall patches are not of significantly lower digestive quality than the pasture average.

Forage intake processes by goats on a massai grass pasture with different sward heights / Processos de ingestão de forragem por caprinos em pastagem com capim-Massai com diferentes Alturas

The objective of this study was to evaluate the forage-intake process of goats feeding on Massai grass pastures with different heights (40, 50, 60, and 70 cm). The experimental design was completely randomized, with treatments corresponding to four sward heights with two replicates over time and space. Collected data were related to the forage-intake process and the chemical and morphological composition of the pasture. Grazing trials (45 min) were performed with four Anglo-Nubian crossbred goats. Total forage mass intake, bite mass, and intake rate were expressed in relation to animal weight. Pasture density, forage mass, and leaf blade increased as forage height increased. The chemical composition of forage at the evaluated heights was similar, except for reduced crude protein content at 70-cm height. Bite rate, intake rate, and time per bite had a quadratic relationship with increasing sward height. The greatest intake rate was observed at 54.7 cm of height, with 0.136 g DM min-1 kg-1 LW. Bite rate exhibited a linear and positive correlation with increase in intake. At the 50-cm height, goats harvested a mass of 3.65 g DM bite-1 kg-1 LW, when they performed 34.5 bites per minute. Adult goats had a greater forage intake on 50-cm high Massai grass pastures because they could obtain a greater bite mass in a shorter time per bite.

Objetivou-se avaliar os processos de ingestão de forragem por caprinos em pastagem com capim-Massai manejado sob diferentes alturas (40, 50, 60 e 70 cm). O delineamento adotado foi o inteiramente casualizado. Os tratamentos corresponderam a quatro alturas do dossel com duas repetições no tempo e no espaço. Os dados coletados foram relacionados aos processos de ingestão de forragem e composição química e morfológica do pasto. Foram realizados testes de pastejo de 45 minutos com quatro cabras mestiças da raça Anglonubiana. A massa de forragem total consumida, massa de bocados e a taxa de ingestão foram expressos em relação ao peso dos animais. Com o incremento da altura da forragem houve aumento na densidade do pasto, massa de forragem e lâmina foliar. A composição química da forragem nas alturas avaliadas foi semelhante, com exceção da redução do teor de proteína bruta aos 70 cm. A taxa de bocado, taxa de ingestão e tempo por bocado apresentou relação quadrática com o aumento da altura. A maior taxa de ingestão foi observada aos 54,7 cm de altura, com 0,136 g MS min-1 kg-1 de PC. A massa de bocado apresentou relação linear e positiva com o aumento da altura. Aos 50 cm as cabras colheram uma massa de 3,65 g MS bocado-1 kg-1 PC, quando realizavam 34,5 bocados por minuto. No pasto de capim-massai aos 50 cm ocorre maior consumo de forragem por cabras adultas, pois os animais conseguem obter uma massa maior do bocado em menor tempo por bocado.

Forage intake processes by goats on a massai grass pasture with different sward heights / Processos de ingestão de forragem por caprinos em pastagem com capim-Massai com diferentes Alturas

The objective of this study was to evaluate the forage-intake process of goats feeding on Massai grass pastures with different heights (40, 50, 60, and 70 cm). The experimental design was completely randomized, with treatments corresponding to four sward heights with two replicates over time and space. Collected data were related to the forage-intake process and the chemical and morphological composition of the pasture. Grazing trials (45 min) were performed with four Anglo-Nubian crossbred goats. Total forage mass intake, bite mass, and intake rate were expressed in relation to animal weight. Pasture density, forage mass, and leaf blade increased as forage height increased. The chemical composition of forage at the evaluated heights was similar, except for reduced crude protein content at 70-cm height. Bite rate, intake rate, and time per bite had a quadratic relationship with increasing sward height. The greatest intake rate was observed at 54.7 cm of height, with 0.136 g DM min-1 kg-1 LW. Bite rate exhibited a linear and positive correlation with increase in intake. At the 50-cm height, goats harvested a mass of 3.65 g DM bite-1 kg-1 LW, when they performed 34.5 bites per minute. Adult goats had a greater forage intake on 50-cm high Massai grass pastures because they could obtain a greater bite mass in a shorter time per bite.(AU)

Objetivou-se avaliar os processos de ingestão de forragem por caprinos em pastagem com capim-Massai manejado sob diferentes alturas (40, 50, 60 e 70 cm). O delineamento adotado foi o inteiramente casualizado. Os tratamentos corresponderam a quatro alturas do dossel com duas repetições no tempo e no espaço. Os dados coletados foram relacionados aos processos de ingestão de forragem e composição química e morfológica do pasto. Foram realizados testes de pastejo de 45 minutos com quatro cabras mestiças da raça Anglonubiana. A massa de forragem total consumida, massa de bocados e a taxa de ingestão foram expressos em relação ao peso dos animais. Com o incremento da altura da forragem houve aumento na densidade do pasto, massa de forragem e lâmina foliar. A composição química da forragem nas alturas avaliadas foi semelhante, com exceção da redução do teor de proteína bruta aos 70 cm. A taxa de bocado, taxa de ingestão e tempo por bocado apresentou relação quadrática com o aumento da altura. A maior taxa de ingestão foi observada aos 54,7 cm de altura, com 0,136 g MS min-1 kg-1 de PC. A massa de bocado apresentou relação linear e positiva com o aumento da altura. Aos 50 cm as cabras colheram uma massa de 3,65 g MS bocado-1 kg-1 PC, quando realizavam 34,5 bocados por minuto. No pasto de capim-massai aos 50 cm ocorre maior consumo de forragem por cabras adultas, pois os animais conseguem obter uma massa maior do bocado em menor tempo por bocado.(AU)

Ingestive behavior of sheep in Panicum and Brachiaria pastures in dry season / Comportamento ingestivo de ovinos em pastos de Panicum E Brachiaria na época seca

The ingestive behavior of sheep supplemented with concentrate in Brachiaria brizantha and Panicum maximum pastures were evaluated during the dry season. Twenty four male sheep of ½ Santa Inês x ½ undefined breed genotypes were used. The treatments were four feeds of cultivar grasses: B. brizantha cvs. Marandu and Piatã, P. maximum cvs. Massai and Aruana. The ingestive behavior of the animals was recorded during two periods of 24 hours. The grazing time, rumination, idleness (minutes day-1), bite rate (bites minute-1), and the frequency of the animals activities in relation to defecation, urination and search for water, concentrated and mineral salt were observed. There was interaction between time of the day and evaluated cultivars based on the grazing time and bite rate response. Sheep grazing on marandu-grass showed higher grazing time between 11 am to 4 pm (257 minutes), compared to animals grazing on aruana-grass (217 minutes). The bite rate was higher for sheep on massai-grass between 11 am to 4 pm (34.32 bits minute- 1). There was no grass effect for idleness and rumination, which can highlight the similarity of chemistry characterization and high percentage of structural components in evaluated pastures. The structural limitations and feed mass composition in the dry season caused effects on the ingestive behavior of idleness and bite rate of supplemented sheep in Brachiaria and Panicum pastures.

Avaliou-se o comportamento ingestivo de ovinos, suplementados com concentrado, em pastos de Brachiaria brizantha e Panicum maximum durante a época seca. Foram utilizados 24 animais, machos inteiros do genótipo ½Santa Inês x ½ sem padrão racial definido. Os tratamentos corresponderam a quatro cultivares de gramíneas forrageiras, sendo B. brizantha cultivares. Marandu e Piatã e P. maximum cultivares. Massai e Aruana. O comportamento ingestivo dos animais foi registrado durante dois períodos de 24 horas. Observou-se o tempo de pastejo, ruminação, ócio (minutos/dia), taxa de bocados (bocados/minuto) e a frequência dos animais em relação à defecação, micção e busca pela água, concentrado e suplemento mineral. Houve interação entre o período do dia e as cultivares avaliadas sobre as respostas de tempo de pastejo e taxa de bocado. Os animais sob pastejo no capim-marandu apresentaram maior tempo de pastejo no período de 11-16h (257 minutos) em relação aos animais do capim-aruana (217 minutos). A taxa de bocados foi maior para os animais do grupo do capim-massai no período 11-16h (34,32 bocados/minuto). Não houve efeito de tratamento para as variáveis de ócio e ruminação, o que pode evidenciar a semelhança da caracterização química e elevada porcentagem de constituintes estruturais nos pastos avaliados. As limitações estruturais, de composição e massa de forragem na época seca provocam efeitos sobre as variáveis de comportamento ingestivo de ovinos suplementados em pastos de Brachiaria e Panicum.

Beef heifers grazing behavior and herbage intake in natural grassland under rotational grazing / Comportamento ingestivo e consumo de forragem de novilhas de corte em pastagem natural manejada com pastoreio rotativo

It was evaluated the effect of two rest intervals between grazing occupations in rotational grazing; 375 and 750 DD (degree-days); based on the cumulative thermal sum necessary for leaf expansion of native grasses of two functional groups over the grazing behavior variables from beef heifers with 12 months old. The experiment was conducted as a completely randomized block design, with two treatments, three replications and measures repeated over time. Grazing behavior was assessed in three occasions (Nov 2011, Jan and Mar 2012), with 24h each. The herbage intake was estimated using an external marker (Cr2O3). The leaf mass was similar among the rest intervals, with a mean of 1261kg DM ha-1. The average grazing time was 627.4min day-1 and bite rate was 37.1 bites min-1. The average number of daily meals was 5.9 with an average of 118.5 minutes. Number of feeding stations visited per minute was 6.1 and the feeding stations permanence time was of 12 seconds. There was a reduction only on grazing time and bite rate over the periods. The range of thermal sums evaluated to define the rest intervals in rotational grazing proved no effect on heifers' grazing behavior and herbage intake.

Foram testados os efeitos de dois intervalos de descanso entre pastoreios, 375 e 750 graus-dia, baseado na soma térmica para a expansão foliar de gramíneas nativas de dois grupos funcionais sobre o comportamento ingestivo de novilhas de corte. O delineamento foi em blocos ao acaso, com dois tratamentos, três repetições de área e medidas repetidas no tempo. Foram realizadas três avaliações (nov/11, jan e mar/12), com 24h cada, utilizando novilhas de corte com 12 meses. Estimou-se o consumo de forragem com marcador externo (Cr2O3). A massa de lâminas foliares foi semelhante entre os tratamentos, apresentando média de 1261kg MS ha-1. O tempo médio dispendido pelos animais em pastejo foi de 627,4 minutos e a taxa de bocados média foi de 37,1 bocados minuto-1. O número médio de refeições foi de 5,9 refeições dia-1, com uma média de 118,5 minutos por refeição. Os animais visitaram, em média, 6,1 estações alimentares por minuto e permaneceram nessas estações por 12 segundos. Houve diferença no tempo de pastejo e taxa de bocado nos períodos, com uma redução ao longo do tempo. Os intervalos de descanso testados provaram ter pouco efeito no comportamento ingestivo e no consumo de forragem das novilhas.

Beef heifers grazing behavior and herbage intake in natural grassland under rotational grazing / Comportamento ingestivo e consumo de forragem de novilhas de corte em pastagem natural manejada com pastoreio rotativo

It was evaluated the effect of two rest intervals between grazing occupations in rotational grazing; 375 and 750 DD (degree-days); based on the cumulative thermal sum necessary for leaf expansion of native grasses of two functional groups over the grazing behavior variables from beef heifers with 12 months old. The experiment was conducted as a completely randomized block design, with two treatments, three replications and measures repeated over time. Grazing behavior was assessed in three occasions (Nov 2011, Jan and Mar 2012), with 24h each. The herbage intake was estimated using an external marker (Cr2O3). The leaf mass was similar among the rest intervals, with a mean of 1261kg DM ha-1. The average grazing time was 627.4min day-1 and bite rate was 37.1 bites min-1. The average number of daily meals was 5.9 with an average of 118.5 minutes. Number of feeding stations visited per minute was 6.1 and the feeding stations permanence time was of 12 seconds. There was a reduction only on grazing time and bite rate over the periods. The range of thermal sums evaluated to define the rest intervals in rotational grazing proved no effect on heifers' grazing behavior and herbage intake.(AU)

Foram testados os efeitos de dois intervalos de descanso entre pastoreios, 375 e 750 graus-dia, baseado na soma térmica para a expansão foliar de gramíneas nativas de dois grupos funcionais sobre o comportamento ingestivo de novilhas de corte. O delineamento foi em blocos ao acaso, com dois tratamentos, três repetições de área e medidas repetidas no tempo. Foram realizadas três avaliações (nov/11, jan e mar/12), com 24h cada, utilizando novilhas de corte com 12 meses. Estimou-se o consumo de forragem com marcador externo (Cr2O3). A massa de lâminas foliares foi semelhante entre os tratamentos, apresentando média de 1261kg MS ha-1. O tempo médio dispendido pelos animais em pastejo foi de 627,4 minutos e a taxa de bocados média foi de 37,1 bocados minuto-1. O número médio de refeições foi de 5,9 refeições dia-1, com uma média de 118,5 minutos por refeição. Os animais visitaram, em média, 6,1 estações alimentares por minuto e permaneceram nessas estações por 12 segundos. Houve diferença no tempo de pastejo e taxa de bocado nos períodos, com uma redução ao longo do tempo. Os intervalos de descanso testados provaram ter pouco efeito no comportamento ingestivo e no consumo de forragem das novilhas.(AU)

Consumo e comportamento ingestivo de cabras em pasto de capim-Marandu / Intake and ingestive behavior of goats on Marandu-grass pasture

O experimento foi conduzido para avaliar a influência da altura do pasto (30, 40, 50 e 60 cm) de capim-Marandu (Brachiaria brizantha) sobre características estruturais do dossel, comportamento em pastejo e processo de ingestão de forragem por caprinos. Para a avaliação do comportamento em pastejo foram utilizadas seis cabras e, do processo de ingestão, quatro cabras, todas da raça Anglonubiana. Adotou-se o delineamento inteiramente casualizado, com duas repetições no espaço e duas repetições no tempo. O aumento na altura do dossel resultou em aumento na massa de forragem e de folhas, colmo, material morto, e densidade de perfílhos, e redução na relação folha/colmo. O tempo de pastejo aumentou e o tempo de ócio diminui com a elevação na altura do pasto. A correlação entre altura do dossel e a profundidade de bocados foi linear positiva (r= 0,99). A massa de forragem consumida, taxa de ingestão e massa do bocado foram maiores aos 60 cm. A correlação entre a altura do pasto e a taxa de bocado foi negativa e, positiva com o tempo por bocado. Em capim-Marandu, maior eficiência de colheita de forragem por caprinos ocorre a 60 cm de altura do dossel.(AU)

The experiment was conducted to evaluate the influence of Marandu-grass (Brachiaria brizantha) pasture height (30, 40, 50 and 60 cm) on the canopy structural traits and grazing behavior and forageingestion process by goats. Six goats were used to evaluate behavior during grazing, and four were used to evaluate the ingestion process - all goats were Anglo-Nubian. The adopted experimental design was completely randomized, with two replicates in space and two replicates in time. Increase in the canopy height resulted in an increase in the masses of forage, leaves, stem, and dead material and tiller density, and reduction in leaf/stem ratio. Grazing time increased and idle time reduced as the canopy height was elevated. The correlation between canopy height and bite depth was positive and linear (r = 0.99). The mass of consumed forage, the intake rate, and the bite mass were higher at 60 cm. The correlation between pasture height and bite rate was negative, whereas the correlation between pasture height and the time per bite was positive. On Marandu-grass pastures, the greatest efficiency in forage harvesting by goats occurs at a canopy height of 60 cm.(AU)

Ingestive behavior and thermoregulatory responses of equine in grazing activities / Comportamento ingestivo e respostas termorregulatórias de equinos em atividades de pastejo

The objective of this article was to evaluate the main physiological responses, as well as the main patterns of ingestive behavior and physiological parameters of equines in activities grazing. Animal behavior is influenced by several factors, such as, climate, temperature, power supply, etc., thus becoming indispensable good management practices, for the animal can play to their best possible performance. The loss of heat in equines that give several ways (conduction, convection, radiation, evaporation), this occurs so that the animal can enter in their zone of thermal comfort (37.5ºC to 38.5ºC).(AU)

O objetivo deste artigo foi avaliar as principais respostas fisiológicas, bem como os principais padrões de comportamento ingestivo e parâmetros fisiológicos de equinos em atividades de pastejo. O comportamento animal recebe influência de diversos fatores, tais como, clima, temperatura, alimentação, etc., tornando-se assim indispensáveis boas práticas de manejo, para que o animal possa expressar o seu melhor desempenho possível. A perda de calor nos equinos se dar de várias maneiras (Condução, convecção, radiação, evaporação), isto ocorre para que o animal possa entrar na sua zona de conforto térmico (37,5°C a 38,5°C).(AU)

Consumo e comportamento ingestivo de cabras em pasto de capim-Marandu / Intake and ingestive behavior of goats on Marandu-grass pasture

O experimento foi conduzido para avaliar a influência da altura do pasto (30, 40, 50 e 60 cm) de capim-Marandu (Brachiaria brizantha) sobre características estruturais do dossel, comportamento em pastejo e processo de ingestão de forragem por caprinos. Para a avaliação do comportamento em pastejo foram utilizadas seis cabras e, do processo de ingestão, quatro cabras, todas da raça Anglonubiana. Adotou-se o delineamento inteiramente casualizado, com duas repetições no espaço e duas repetições no tempo. O aumento na altura do dossel resultou em aumento na massa de forragem e de folhas, colmo, material morto, e densidade de perfílhos, e redução na relação folha/colmo. O tempo de pastejo aumentou e o tempo de ócio diminui com a elevação na altura do pasto. A correlação entre altura do dossel e a profundidade de bocados foi linear positiva (r= 0,99). A massa de forragem consumida, taxa de ingestão e massa do bocado foram maiores aos 60 cm. A correlação entre a altura do pasto e a taxa de bocado foi negativa e, positiva com o tempo por bocado. Em capim-Marandu, maior eficiência de colheita de forragem por caprinos ocorre a 60 cm de altura do dossel.

The experiment was conducted to evaluate the influence of Marandu-grass (Brachiaria brizantha) pasture height (30, 40, 50 and 60 cm) on the canopy structural traits and grazing behavior and forageingestion process by goats. Six goats were used to evaluate behavior during grazing, and four were used to evaluate the ingestion process - all goats were Anglo-Nubian. The adopted experimental design was completely randomized, with two replicates in space and two replicates in time. Increase in the canopy height resulted in an increase in the masses of forage, leaves, stem, and dead material and tiller density, and reduction in leaf/stem ratio. Grazing time increased and idle time reduced as the canopy height was elevated. The correlation between canopy height and bite depth was positive and linear (r = 0.99). The mass of consumed forage, the intake rate, and the bite mass were higher at 60 cm. The correlation between pasture height and bite rate was negative, whereas the correlation between pasture height and the time per bite was positive. On Marandu-grass pastures, the greatest efficiency in forage harvesting by goats occurs at a canopy height of 60 cm.

Ingestive behavior and thermoregulatory responses of equine in grazing activities / Comportamento ingestivo e respostas termorregulatórias de equinos em atividades de pastejo

The objective of this article was to evaluate the main physiological responses, as well as the main patterns of ingestive behavior and physiological parameters of equines in activities grazing. Animal behavior is influenced by several factors, such as, climate, temperature, power supply, etc., thus becoming indispensable good management practices, for the animal can play to their best possible performance. The loss of heat in equines that give several ways (conduction, convection, radiation, evaporation), this occurs so that the animal can enter in their zone of thermal comfort (37.5ºC to 38.5ºC).

O objetivo deste artigo foi avaliar as principais respostas fisiológicas, bem como os principais padrões de comportamento ingestivo e parâmetros fisiológicos de equinos em atividades de pastejo. O comportamento animal recebe influência de diversos fatores, tais como, clima, temperatura, alimentação, etc., tornando-se assim indispensáveis boas práticas de manejo, para que o animal possa expressar o seu melhor desempenho possível. A perda de calor nos equinos se dar de várias maneiras (Condução, convecção, radiação, evaporação), isto ocorre para que o animal possa entrar na sua zona de conforto térmico (37,5°C a 38,5°C).

Diurnal behavior of dairy cows on alfalfa pastures supplemented with corn silage and concentrate / Comportamento diurno de vacas leiteiras em pastagem de alfafa suplementada com silagem de milho e concentrado

The diurnal ingestive behavior of dairy cows on alfalfa pastures supplemented with corn silage and concentrate was evaluated. Sixteen Holstein dairy cows in mid-lactation were used in a randomized block design. The treatments were partial replacement of corn silage for limited alfalfa grazing time (LGT) or unlimited grazing time (UGT). Rotational alfalfa grazing was used and the amount of concentrate fed was the same in both treatments. Behavioral activities observed were time spent grazing, rumination and idleness, during four days, from 8:00 AM to 7:00 PM. From 2:00 to 5:00 PM grazing was intermittent and short-lasting on UGT, whereas on LGT grazing activity was longer than UGT in the morning and at the end of the afternoon. Restriction in available grazing time clearly influenced the distribution of time spent with grazing, rumination and idleness throughout the day. However, this effect was not observed in the total time spent in these activities...

Avaliou-se o comportamento ingestivo de vacas leiteiras com acesso restrito ou irrestrito a uma pastagem de alfafa suplementada com silagem de milho e concentrado. Utilizaram-se 16 vacas da raça Holandesa, em estádio médio de lactação, em delineamento em blocos ao acaso. Os tratamentos foram: silagem de milho substituída parcialmente por pastejo restrito ou irrestrito de alfafa. As vacas foram manejadas em sistema de pastejo rotacionado e a quantidade de concentrado foi igual em ambos os tratamentos. As atividades comportamentais observadas foram pastejo, ruminação e ócio, das oito às 19 horas, em quatro ocasiões. No pastejo irrestrito, observou-se que, nos horários entre 14 e 17 horas, o pastejo foi intermitente e de curta duração, enquanto no pastejo restrito a atividade de pastejo foi mais prolongada durante a manhã e no final da tarde. A restrição no tempo disponível para pastejo da alfafa influenciou marcadamente a distribuição, no decorrer do dia, do tempo gasto em pastejo, da ruminação e do ócio. Entretanto, não se observou este efeito no tempo total gasto nestas atividades...

Diurnal behavior of dairy cows on alfalfa pastures supplemented with corn silage and concentrate / Comportamento diurno de vacas leiteiras em pastagem de alfafa suplementada com silagem de milho e concentrado

The diurnal ingestive behavior of dairy cows on alfalfa pastures supplemented with corn silage and concentrate was evaluated. Sixteen Holstein dairy cows in mid-lactation were used in a randomized block design. The treatments were partial replacement of corn silage for limited alfalfa grazing time (LGT) or unlimited grazing time (UGT). Rotational alfalfa grazing was used and the amount of concentrate fed was the same in both treatments. Behavioral activities observed were time spent grazing, rumination and idleness, during four days, from 8:00 AM to 7:00 PM. From 2:00 to 5:00 PM grazing was intermittent and short-lasting on UGT, whereas on LGT grazing activity was longer than UGT in the morning and at the end of the afternoon. Restriction in available grazing time clearly influenced the distribution of time spent with grazing, rumination and idleness throughout the day. However, this effect was not observed in the total time spent in these activities.(AU)

Avaliou-se o comportamento ingestivo de vacas leiteiras com acesso restrito ou irrestrito a uma pastagem de alfafa suplementada com silagem de milho e concentrado. Utilizaram-se 16 vacas da raça Holandesa, em estádio médio de lactação, em delineamento em blocos ao acaso. Os tratamentos foram: silagem de milho substituída parcialmente por pastejo restrito ou irrestrito de alfafa. As vacas foram manejadas em sistema de pastejo rotacionado e a quantidade de concentrado foi igual em ambos os tratamentos. As atividades comportamentais observadas foram pastejo, ruminação e ócio, das oito às 19 horas, em quatro ocasiões. No pastejo irrestrito, observou-se que, nos horários entre 14 e 17 horas, o pastejo foi intermitente e de curta duração, enquanto no pastejo restrito a atividade de pastejo foi mais prolongada durante a manhã e no final da tarde. A restrição no tempo disponível para pastejo da alfafa influenciou marcadamente a distribuição, no decorrer do dia, do tempo gasto em pastejo, da ruminação e do ócio. Entretanto, não se observou este efeito no tempo total gasto nestas atividades.(AU)

Ingestive behavior of hoggets given different types of supplement on ryegrass pasture / Comportamento ingestivo de borregas recebendo diferentes tipos de suplemento em pastagem de azevém

The ingestive behavior of hoggets was assessed under intermittent grazing method with three days of paddock occupation. These animals remained exclusively on Italian ryegrass (Lolium multiflorum Lam) pasture or on ryegrass pasture and receiving 1.2% of DM in relation to body weight of rice paddy, corn grain or whole rice bran as a supplement during the vegetative, pre-flowering and flowering phenological stages of ryegrass. The experimental design was a randomized split-split plots wherein the types of supplement were considered as main plot, phenological stages as subplot and the days of occupation as sub-subplots. The hoggets grazed for a longer time when kept exclusively on ryegrass pasture. The hoggets grazed for less time in the vegetative stage of ryegrass and on the third day of paddock occupation. Feeding behavior of hoggets is changed when supplementation is provided in different phenological stages of the grass and days of paddock occupation.(AU)

Em método intermitente de pastejo, com três dias de ocupação dos piquetes, foi avaliado o comportamento ingestivo de borregas. Esses animais permaneceram em pastagem exclusiva de azevém (Lolium multiflorum Lam.) ou em pastagem de azevém recebendo 1,2% de MS em relação ao peso corporal de grão de arroz em casca, grão de milho moído ou farelo de arroz integral como suplemento durante os estádios fenológicos vegetativo, pré-florescimento e florescimento do azevém. O delineamento experimental foi o inteiramente casualizado em parcelas sub-subdivididas, sendo considerados os tipos de suplemento como parcelas principais, os estádios fenológicos como subparcelas e os dias de ocupação como sub-subparcelas. As borregas pastejaram por maior tempo quando permaneceram exclusivamente em azevém. As borregas pastejaram por menor tempo no estádio vegetativo do azevém e no terceiro dia de ocupação dos piquetes. O comportamento ingestivo de borregas é alterado quando é fornecido suplemento nos diferentes estádios fenológicos do pasto e nos dias de ocupação dos piquetes.(AU)

Ingestive behavior of hoggets given different types of supplement on ryegrass pasture / Comportamento ingestivo de borregas recebendo diferentes tipos de suplemento em pastagem de azevém

The ingestive behavior of hoggets was assessed under intermittent grazing method with three days of paddock occupation. These animals remained exclusively on Italian ryegrass (Lolium multiflorum Lam) pasture or on ryegrass pasture and receiving 1.2% of DM in relation to body weight of rice paddy, corn grain or whole rice bran as a supplement during the vegetative, pre-flowering and flowering phenological stages of ryegrass. The experimental design was a randomized split-split plots wherein the types of supplement were considered as main plot, phenological stages as subplot and the days of occupation as sub-subplots. The hoggets grazed for a longer time when kept exclusively on ryegrass pasture. The hoggets grazed for less time in the vegetative stage of ryegrass and on the third day of paddock occupation. Feeding behavior of hoggets is changed when supplementation is provided in different phenological stages of the grass and days of paddock occupation.

Em método intermitente de pastejo, com três dias de ocupação dos piquetes, foi avaliado o comportamento ingestivo de borregas. Esses animais permaneceram em pastagem exclusiva de azevém (Lolium multiflorum Lam.) ou em pastagem de azevém recebendo 1,2% de MS em relação ao peso corporal de grão de arroz em casca, grão de milho moído ou farelo de arroz integral como suplemento durante os estádios fenológicos vegetativo, pré-florescimento e florescimento do azevém. O delineamento experimental foi o inteiramente casualizado em parcelas sub-subdivididas, sendo considerados os tipos de suplemento como parcelas principais, os estádios fenológicos como subparcelas e os dias de ocupação como sub-subparcelas. As borregas pastejaram por maior tempo quando permaneceram exclusivamente em azevém. As borregas pastejaram por menor tempo no estádio vegetativo do azevém e no terceiro dia de ocupação dos piquetes. O comportamento ingestivo de borregas é alterado quando é fornecido suplemento nos diferentes estádios fenológicos do pasto e nos dias de ocupação dos piquetes.