Review of the bicolora Fang and binghami Hampson species-groups of the genus Ovipennis Hampson with descriptions of five new species from Southwestern China and Indochina (Lepidoptera, Erebidae, Arctiinae, Lithosiini).

The Ovipennis bicolora Fang, 1986 and Ovipennis binghami Hampson, 1903 species-groups are reviewed. The identities of the records of O.binghami reported outside of its type locality are clarified. Ovipennis bicolora is rediscovered with the male and female genitalia illustrated for the first time. The male and female genitalia of O.thomasi ern, 2009 are illustrated for the first time and its specific status is confirmed. Five new species are described: O.hanae S.-Y. Huang, Volynkin & ern, sp. n. (Southwestern China), O.regina Volynkin, ern, S.-Y. Huang & Saldaitis, sp. n. (Northern Thailand and Southwestern China), O.takia Volynkin, S.-Y. Huang & ern, sp. n. (Western and Central Thailand), O.kitchingi Volynkin, S.-Y. Huang & ern, sp. n. (Northern Thailand), and O.sapa Volynkin, S.-Y. Huang, ern & Saldaitis, sp. n. (Northern Vietnam). Adults and genitalia of the new species are illustrated and compared with its congeners.

Letrasilta, a new genus for the Striatella cernyi Volynkin species-group with description of a new species from Xizang, China, and a replacement name for the genus group name Striatella Volynkin & S.-Y. Huang, 2019 (Lepidoptera: Erebidae: Arctiinae: Lithosiini).

The name Striatochrista nom. n. is introduced as replacement for Striatella Volynkin & S.-Y. Huang, 2019. A new genus, Letrasilta S.-Y. Huang & Volynkin gen. n. is erected to include the Striatella cernyi species-group with the new species, L. ratnasambhava S.-Y. Huang, Volynkin & Yin sp. n. from Xizang, southwestern China designated as the type species. Based on the molecular phylogenetic analysis, the new genus is found to be sister to the clade (Aberrasine + ((Indiania + Idopterum) + Striatochrista nom. n.)) but is distinguished from all the relevant genera by the unique genitalia features. Letrasilta cernyi (Volynkin, 2018) comb. n. is also reported from India for the first time. Adults and genitalia of the aforementioned taxa are illustrated. A checklist of the genus Striatochrista is also provided.

Taxonomic review of the genus Tarika Moore with descriptions of six new species from Nepal, China and Indochina (Lepidoptera: Erebidae: Arctiinae).

The Asiatic footman moth genus Tarika Moore, 1878, which used to be considered as monotypic is reviewed. Six new species are described: T. annapurna sp. n. (Сentral Nepal), T. erlanga sp. n. (China: Sichuan), T. reducta sp. n. (North Thailand), T. danieli sp. n. (China: Yunnan, Sichuan, Fujian, Shandong), T. kinha sp. n. (North Vietnam and China: Yunnan), and T. biserratula sp. n. (China: Hunan). Lectotypes are designated for Lithosia varana Moore, 1865 and Tarika nivea Moore, 1878. Adults, male and female genitalia are illustrated.

A new species of Karschiola Gaede from Mozambique and Zimbabwe (Lepidoptera: Erebidae: Arctiinae) with updated information on the distribution of the genus.

The present paper provides the description of a new species of the genus Karschiola Gaede, 1926 from central Mozambique and Zimbabwe: K. ndzou sp. n. The pairwise genetic distance between the two Karschiola species is calculated and a neighbour-joining tree based on DNA barcodes of two K. holoclera and five K. ndzou sp. n. specimens is provided. The paper is illustrated with eight colour photos of adults and 18 genitalia images, a distribution map and two habitat photos.

Taxonomic review of the Oriental genus Gandhara Moore with descriptions of four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini).

The genus Gandhara Moore, 1878 is reviewed. Four new species are described: Gandhara typica Volynkin & ern, sp. n. (Nepal, northeast India, northern Myanmar and northern Thailand), Gandhara clava Volynkin & ern, sp. n. (Laos and northern Thailand), Gandhara emgai Volynkin & ern, sp. n. (northern Vietnam), and Gandhara interrogativa Volynkin, ern, S.-Y. Huang & H.-L. Hu, sp. n. (northern Vietnam and southern China). Gandhara typica sp. n. is fixed as the type species of the genus Gandhara whereas Lithosia serva Walker, 1854 considered by authors as the type species of Gandhara by misidentification is transferred to the genus Collita Moore, 1878. Two new combinations are introduced: Gandhara conica (Fang, 2000), comb. n. and Collita serva (Walker, 1854), comb. n. A lectotype is designated for Lithosia serva Walker, 1854 in order to stabilise the nomenclature. Adults as well as male and female genitalia are illustrated.

On the taxonomy of the genus Isochlora Staudinger with descriptions of two new species from Mongolia and Qinghai, China (Lepidoptera: Noctuidae: Noctuinae).

Two new species of the genus Isochlora Staudinger, 1882 are described: I. hreblayi Volynkin, Tth, Titov & Saldaitis, sp. n. (western Mongolia) and I. kozlovi Volynkin, Titov, Matov & Saldaitis, sp. n. (Qinghai Province, China). The type species of the genus-group names Chamyla Staudinger, 1900 and Grumia Alphraky, 1892 (I. arctomys Alphraky, 1897 and I. flora (Alphraky, 1892), respectively) are examined, and their synonymy with Isochlora is revised. The synonymy of Chamyla idia Staudinger, 1900 with Isochlora arctomys Alphraky, 1897 is revised as junior synonymies. Lectotypes are designated for Isochlora arctomys Alphraky, 1897, Chamyla idia Staudinger, 1900 and Grumia flora Alphraky, 1892. Isochlora intricans (Alphraky, 1882) is reported from Kazakhstan for the first time. Adults as well as male and female genitalia are illustrated.

Taxonomic review of the Cyana adelina (Staudinger) species-group (Lepidoptera: Erebidae: Arctiinae) with descriptions of three new species and one new subspecies from Laos, Thailand and China.

The Cyana adelina (Staudinger, 1887) species-group is established and reviewed. Three new species and one new subspecies are described: Cyana yunnanensis regia ssp. n. (Northern Thailand and northern Laos), Cyana dubernardi sp. n. (China: Yunnan), Cyana khami sp. n. (central Laos), and Cyana karsticola sp. n. (northern Laos). Two new synonyms are established: Cyana yunnanensis yunnanensis (Hampson, 1903) (Chionaema) = Chionaema hoenei Daniel, 1952, syn. n., and Cyana griseilinea de Joannis, 1930 = Cyana succincta ern, 2016, syn. n. Lectotypes are designated for Bizone adelina Staudinger, 1887 and Bizone pratti Elwes, 1890. Adults, male and female genitalia are illustrated.

Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus HebenaWalker, 1856, with descriptions of six new species and one new subspecies(Lepidoptera: Erebidae: Arctiinae: Syntomini).

A revision of the subgenus Hebena Walker, 1856 of the syntomine genus Metarctia Walker, 1855 is given based on integrative taxonomic analyses. Six new species (Metarctia (Hebena) manfredi sp. n., M. (H.) elleni sp. n., M. (H.) brigitta sp. n., M. (H). lukaszi sp. n., M. (H.) smithi sp. n., and M. (H.) haraldsulaki sp.n.) and one new subspecies (M. (H.) smithi transvallesiana ssp. n.) are described. Metarctia (H.) kelleni (Snellen, 1886) stat. rev. is reinstated from synonymy with M. (H.) rubra (Walker, 1856) and Metarctia (H.) subincarnata Kiriakoff, 1954 syn. n. is synonymised with M. (H.) henrardi Kiriakoff, 1953 and Metarctia cinnamomea (Wallengren, 1860) syn. n. with M. (H.) rubra (Walker, 1856). DNA barcodes were obtained for 116 specimens representing 7 taxa, and genetic analyses were performed using Maximum Likelihood and Bayesian Inference approaches; a DNA barcode tree resulting from the latter is illustrated. Pairwise distances of barcodes between taxa are provided where available. The adults and genitalia of all taxa, their habitats and distribution are illustrated in 19 colour plates and 5 distribution maps.

On the identity of Conisania leuconephra Draudt, 1950 with a description of a new species from Xizang, China (Lepidoptera: Noctuidae: Noctuinae).

The identity of Conisania leuconephra (Draudt, 1950) from Yunnan (China) is revised. A new species Conisania sejilaensis sp. nov. is described from Xizang (China). The adults as well as the male and the female genitalia of the new and related species are illustrated.

On the taxonomy of Tigrioides leucanioides (Walker) with descriptions of two new genera and ten new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae: Lithosiini).

The generic placement and the taxonomic structure of the complex traditionally identified in the collections as Tigrioides leucanioides (Walker, 1862) are revised. Two new genera and ten new species are described: Leucanithosia gen. n., Leucanithosia diehli sp. n. (Indonesia: northern Sumatra Island), Leucanithosia paraliaka sp. n. (Indonesia: northern Sumatra Island), Leucanithosia nansabon sp. n. (northern Myanmar), Leucanithosia witti sp. n. (Thailand, Laos, Southwest China (Yunnan) and Northeast India), Leucanithosia leptivalva sp. n. (the Philippines: Palawan Island), Leucanithosia inusitata sp. n. (Indonesia: northern Sumatra Island), Leucanithosia lampunga sp. n. (Indonesia: southern Sumatra Island), Philippinithosia gen. n., Philippinithosia lourensi sp. n. (the Philippines: Luzon, Negros and Mindoro Islands), Philippinithosia settelei sp. n. (the Philippines: Mindanao Island), and Philippinithosia reducta sp. n. (the Philippines: Mindanao, Leyte and Samar Islands). A new combination is established: Leucanithosia leucanioides (Walker, 1862), comb. n. The lectotype is designated for Lithosia leucanioides Walker, 1862. Adults, male and female genitalia are illustrated.

Contribution to the knowledge of the genus Nephelomilta Hampson, 1900(Lepidoptera, Erebidae, Arctiinae) from China with descriptions of two new species.

New information on the genus Nephelomilta Hampson, 1900 from China is reported. Two new species are described from southwestern China: N. medogensis Huang, Volynkin & Saldaitis, sp. n. from Motuo County, Xizang Autonomous Region and N. nu Huang, Volynkin & Saldaitis, sp. n. from Nujiang Valley, Yunnan Province. The female of N. yasunorikishidai Huang & Wang, 2019 is discovered and illustrated for the first time, and the species is erected into a separate species-group. Adults and genitalia of the aforementioned taxa and related ones as well are illustrated.

On the taxonomy of the genus Teulisna Walker from India with a description of a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini).

The taxonomy of two Indian species of the genus Teulisna Walker, 1862, T. basigera (Walker, [1865]) and T. inducta (Walker, [1865]), is revised. One species is reinstated from the synonymy with T. basigera: Teulisna ruma (Swinhoe, [1890] 1889), stat. rev. A new species, Teulisna munnara sp. n. is described from the state of Kerala (South India). Lectotypes are designated for Cossa basigera Walker, [1865] 1864, Lithosia inducta Walker, [1865] 1864, and Cossa ruma Swinhoe, [1890] 1889. Adults as well as male and female genitalia of all the species considered are illustrated.

Redescription of the genus Arcilasisa Walker with notes on its taxonomic placement (Lepidoptera: Noctuidae: Noctuinae).

The poorly known monotypic noctuid genus Arcilasisa Walker, 1865 is redescribed. The hitherto unknown male and female genitalia of the type species of the genus, A. sobria Walker, 1865 are illustrated and diagnosed for the first time. The genus is placed in the Apamea Ochsenheimer, 1816 generic complex belonging to the subtribe Apameina Guene of the tribe Apameini Guene. The lectotype for the type species of the genus, Arcilasisa sobria Walker, 1865 is designated.

Three new species of Miltochrista Hbner from India, Myanmar, Nepal and China (Lepidoptera: Erebidae: Arctiinae: Lithosiini).

Three new species of the genus Miltochrista Hbner, [1819] are described: M. idiomorfa sp. n. (India, Nepal, N Myanmar and SW China), M. dankana sp. n. (India and north-western Myanmar) and M. etalina sp. n. (Northeast India). Miltochrista straminea Walker, 1856, stat. rev. is restored to species level. The lectotype for Miltochrista straminea is designated. Adults, male and female genitalia of the new and similar species are illustrated.

Anchoscelis fuscomixta noravanka, a new subspecies from Armenia (Lepidoptera: Noctuidae: Noctuinae).

A new subspecies of the genus Anchoscelis Guene, 1839, Anchoscelis fuscomixta noravanka ssp. n. is described from south-west Armenia. The new subspecies belongs to the nominate subgenus of Anchoscelis and is similar, besides Anchoscelis fuscomixta (Gyulai L. Ronkay, 2006), to Anchoscelis pauli (Staudinger, 1892), Anchoscelis kindermanni (Fischer von Rslerstamm, 1839) and Anchoscelis consueta (Herrich-Schffer, 1852). Adults and male and female genitalia of the new subspecies and the similar taxa are illustrated. The habitat of the new subspecies is characterised and illustrated.

On the taxonomy of Barsine striata (Bremer amp; Grey) and Barsine pulchra (Butler) (Lepidoptera: Erebidae: Arctiinae) with descriptions of three new species from Indochina.

The taxonomy of Barsine striata (Bremer Grey, 1852) and B. pulchra (Butler, 1877) is revised. Three new species are described: B. evrys sp. n. (Central and South Vietnam, and South-East Thailand), B. robusta sp. n. (North Thailand), and B. laotiana sp. n. (entral Laos). One taxon is revived from synonymy and upgraded to subspecies level: Barsine pulchra pulcherrima (Staudinger, 1887), stat. rev. Miltochrista gratiosa kurilensis Bryk, 1942, is excluded from the synonymy with Barsine striata and placed in synonymy with B. pulchra (=Miltochrista gratiosa kurilensis Bryk, 1942, stat. rev.). Lectotypes for Lithosia striata Bremer Grey, 1852, Hypoprepia lanceolata Walker, 1854 and Miltochrista pulchra Butler, 1877 are designated. Adults, male and female genitalia are illustrated.

Two remarkable new species of the genus Barsine Walker, 1854 from China (Lepidoptera: Erebidae: Arctiinae: Lithosiini).

Two remarkable new species of the genus Barsine Walker, 1854 are described: Barsine qinxiae S.-Y. Huang Volynkin sp. n. from Zhejiang, Eastern China, and B. jingkuni S.-Y. Huang Volynkin sp. n. from Sichuan, Western China. The diagnostic comparison is made with Barsine roseata (Walker, 1864), B. hoenei (Reich, 1937) and B. pardalis (Mell, 1922). The lectotype for Miltochrista hoenei Reich, 1937 is designated. Adults and genitalia of the aforementioned taxa are illustrated.

Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera: Erebidae: Arctiinae).

The Manulea tienmushanica (Daniel, 1954) species group is established, characterised and reviewed. Two new species are described: Manulea (M.) salweena sp. n. (China: Yunnan), and Manulea (M.) diaforetica sp. n. (China: Yunnan). Two new combinations are established: Manulea (M.) tienmushanica (Daniel, 1954), comb. n. and Manulea (M.) nigripuncta (Fang, 2000), comb. n. Adults, male and female genitalia are illustrated.

Contribution to the knowledge of the genus Ovipennis Hampson, 1900 with descriptions of two new species from Southern and Southwestern China (Lepidoptera, Erebidae, Arctiinae, Lithosiini).

The knowledge of the genus Ovipennis Hampson, 1900 is updated. Nanarsine milani (ern, 2009) is recorded from China and Vietnam for the first time and transferred to the genus Ovipennis based on multilocus molecular phylogenetic and morphological analyses: O. milani (ern, 2009), comb. n. Two new species related to Ovipennis semilutea species-group are described: O. mixta Huang, Zhao, Volynkin Han, sp. n. from Guizhou and Hunan and O. multicornuta Huang, Volynkin, Zhao Han, sp. n. from Sichuan. Adults and genitalia of the aforementioned taxa are illustrated.

Two new species of the subgenus Cryptanaema de Vos of the genus Cyana Walker from eastern Indonesia (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Nudariina).

Two new species of the subgenus Cryptanaema de Vos, 2017 of the genus Cyana Walker, 1854 are described from eastern Indonesia: Cyana (Cryptanaema) ngata sp. n. (Western New Guinea) and Cyana (Cryptanaema) sibela sp. n. (Maluku Islands). Adults, male and female genitalia of the new and similar species are illustrated.