Escarpment evolution drives the diversification of the Madagascar flora.

Madagascar exhibits high endemic biodiversity that has evolved with sustained and stable rates of speciation over the past several tens of millions of years. The topography of Madagascar is dominated by a mountainous continental rift escarpment, with the highest plant diversity and rarity found along the steep, eastern side of this geographic feature. Using a process-explicit model, we show that precipitation-driven erosion and landward retreat of this high-relief topography creates transient habitat organization through multiple mechanisms, including catchment expansion, isolation of highland remnants, and formation of topographic barriers. Habitat isolation and reconnection on a million-year timescale serves as an allopatric speciation pump creating the observed biodiversity.

Phytodiversity is associated with habitat heterogeneity from Eurasia to the Hengduan Mountains.

The geographic distribution of plant diversity matches the gradient of habitat heterogeneity from lowlands to mountain regions. However, little is known about how much this relationship is conserved across scales. Using the World Checklist of Vascular Plants and high-resolution biodiversity maps developed by species distribution models, we investigated the associations between species richness and habitat heterogeneity at the scales of Eurasia and the Hengduan Mountains (HDM) in China. Habitat heterogeneity explains seed plant species richness across Eurasia, but the plant species richness of 41/97 HDM families is even higher than expected from fitted statistical relationships. A habitat heterogeneity index combining growing degree days, site water balance, and bedrock type performs better than heterogeneity based on single variables in explaining species richness. In the HDM, the association between heterogeneity and species richness is stronger at larger scales. Our findings suggest that high environmental heterogeneity provides suitable conditions for the diversification of lineages in the HDM. Nevertheless, habitat heterogeneity alone cannot fully explain the distribution of species richness in the HDM, especially in the western HDM, and complementary mechanisms, such as the complex geological history of the region, may have contributed to shaping this exceptional biodiversity hotspot.

Greater Alpine river network evolution, interpretations based on novel drainage analysis.

The Central European drainage system is dominated by four major rivers (the Danube, Rhine, Rhône and Po). The geometry of these drainage basins has evolved through the history of Alpine and Carpathian orogeny. Analysis of the modern river geometry reveals the geometric stability or instability of the drainage network and enables interpretation of the erosion and exhumation pattern. We characterize the river basin geometry and inter-basin relief through metrics including the quantity χ that relates to the catchment area, and a catchment restricted minimum elevation (CRM) metric. The interpretations from the maps are in agreement with known river captures and morphological features. The χ-map reveals additional systematic, large-scale transients, consistent with ongoing basin changes, mainly manifesting in the Danube losing catchment area. We postulate that the changes are related to the longitudinal initiation of the Danube River in the Alpine foreland, augmented by the formation of the Carpathians and the filling of the Pannonian Basin, which resulted in an elongation of the Danube river basin. We conclude that the Danube has lacked erosional power throughout its history and therefore been victim to capture and area loss.

Transience of the North American High Plains landscape and its impact on surface water.

Ecosystem diversity and human activity in dry climates depend not just on the magnitude of rainfall, but also on the landscape's ability to retain water. This is illustrated dramatically in the High Plains of North America, where despite the semi-arid modern and past climate, the hydrologic conditions are diverse. Large rivers sourced in the Rocky Mountains cut through elevated plains that exhibit limited river drainage but widespread surface water in the form of ephemeral (seasonal) playa lakes1, as well as extensive groundwater hosted in the High Plains aquifer of the Ogallala formations2. Here we present a model, with supporting evidence, which shows that the High Plains landscape is currently in a transient state, in which the landscape has bifurcated into an older region with an inefficient river network and a younger, more efficient, river channel network that is progressively cannibalizing the older region. The older landscape represents the remnants of the Ogallala sediments that once covered the entirety of the High Plains, forming depositional fans that buried the pre-existing river network and effectively 'repaved' the High Plains3-6. Today we are witnessing the establishment of a new river network that is dissecting the landscape, capturing channels and eroding these sediment fans. Through quantitative analysis of the geometry of the river network, we show how network reorganization has resulted in a distinctive pattern of erosion, whereby the largest rivers have incised the older surface, removed fan heads near the Rocky Mountains and eroded the fan toes, but left portions of the central fan surface and the Ogallala sediments largely intact. These preserved fan surfaces have poor surface water drainage, and thus retain ephemeral water for wetlands and groundwater recharge. Our findings suggest that the surface hydrology and associated ecosystems are transient features on million-year timescales, and reflect the mode of landscape evolution.

A global dataset of river network geometry.

The plan-form structure of the world's river basins contains extensive information regarding tectonic, paleo-geographic and paleo-climate conditions, but interpretation of this structure is complicated by the need to disentangle these processes from the autogenic behavior of fluvial processes. One method of interpreting this structure is by integrating channel length and drainage area as characterized by the scaling relationship between slope and area, resulting in a characteristic length parameter, referred to in recent studies as χ. In this paper, we apply this methodology at a continental scale by calculating χ for the world's river networks. Mapping of χ', a modified version of χ including the influence of precipitation distribution on river discharge and correction of base level for χ' in closed basins, illustrates the geometric structure of global river networks, thus highlighting where tectonics or changing climate have resulted in an apparent disequilibrium of the river channel geometry. Our global χ maps quantify a dynamic view of Earth's river networks and help to identify past and ongoing evolution of Earth's landscape.

In situ low-relief landscape formation as a result of river network disruption.

Landscapes on Earth retain a record of the tectonic, environmental and climatic history under which they formed. Landscapes tend towards an equilibrium in which rivers attain a stable grade that balances the tectonic production of elevation and with hillslopes that attain a gradient steep enough to transport material to river channels. Equilibrium low-relief surfaces are typically found at low elevations, graded to sea level. However, there are many examples of high-elevation, low-relief surfaces, often referred to as relict landscapes, or as elevated peneplains. These do not grade to sea level and are typically interpreted as uplifted old landscapes, preserving former, more moderate tectonic conditions. Here we test this model of landscape evolution through digital topographic analysis of a set of purportedly relict landscapes on the southeastern margin of the Tibetan Plateau, one of the most geographically complex, climatically varied and biologically diverse regions of the world. We find that, in contrast to theory, the purported surfaces are not consistent with progressive establishment of a new, steeper, river grade, and therefore they cannot necessarily be interpreted as a remnant of an old, low relief surface. We propose an alternative model, supported by numerical experiments, in which tectonic deformation has disrupted the regional river network, leaving remnants of it isolated and starved of drainage area and thus unable to balance tectonic uplift. The implication is that the state of low relief with low erosion rate is developing in situ, rather than preserving past erosional conditions.

Dynamic reorganization of river basins.

River networks evolve as migrating drainage divides reshape river basins and change network topology by capture of river channels. We demonstrate that a characteristic metric of river network geometry gauges the horizontal motion of drainage divides. Assessing this metric throughout a landscape maps the dynamic states of entire river networks, revealing diverse conditions: Drainage divides in the Loess Plateau of China appear stationary; the young topography of Taiwan has migrating divides driving adjustment of major basins; and rivers draining the ancient landscape of the southeastern United States are reorganizing in response to escarpment retreat and coastal advance. The ability to measure the dynamic reorganization of river basins presents opportunities to examine landscape-scale interactions among tectonics, erosion, and ecology.

Worldwide acceleration of mountain erosion under a cooling climate.

Climate influences the erosion processes acting at the Earth's surface. However, the effect of cooling during the Late Cenozoic era, including the onset of Pliocene-Pleistocene Northern Hemisphere glaciation (about two to three million years ago), on global erosion rates remains unclear. The uncertainty arises mainly from a lack of consensus on the use of the sedimentary record as a proxy for erosion and the difficulty of isolating the respective contributions of tectonics and climate to erosion. Here we compile 18,000 bedrock thermochronometric ages from around the world and use a formal inversion procedure to estimate temporal and spatial variations in erosion rates. This allows for the quantification of erosion for the source areas that ultimately produce the sediment record on a timescale of millions of years. We find that mountain erosion rates have increased since about six million years ago and most rapidly since two million years ago. The increase of erosion rates is observed at all latitudes, but is most pronounced in glaciated mountain ranges, indicating that glacial processes played an important part. Because mountains represent a considerable fraction of the global production of sediments, our results imply an increase in sediment flux at a global scale that coincides closely with enhanced cooling during the Pliocene and Pleistocene epochs.

Links between erosion, runoff variability and seismicity in the Taiwan orogen.

The erosion of mountain belts controls their topographic and structural evolution and is the main source of sediment delivered to the oceans. Mountain erosion rates have been estimated from current relief and precipitation, but a more complete evaluation of the controls on erosion rates requires detailed measurements across a range of timescales. Here we report erosion rates in the Taiwan mountains estimated from modern river sediment loads, Holocene river incision and thermochronometry on a million-year scale. Estimated erosion rates within the actively deforming mountains are high (3-6 mm yr(-1)) on all timescales, but the pattern of erosion has changed over time in response to the migration of localized tectonic deformation. Modern, decadal-scale erosion rates correlate with historical seismicity and storm-driven runoff variability. The highest erosion rates are found where rapid deformation, high storm frequency and weak substrates coincide, despite low topographic relief.